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The northern brown argus (''Aricia artaxerxes'') is a
butterfly Butterflies are insects in the macrolepidopteran clade Rhopalocera from the Order (biology), order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The ...
in the family
Lycaenidae Lycaenidae is the second-largest family of butterflies (behind Nymphalidae, brush-footed butterflies), with over 6,000 species worldwide, whose members are also called gossamer-winged butterflies. They constitute about 30% of the known butterfl ...
. It is found throughout much of the
Palearctic realm The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa. The realm consists of several bioregions: the Euro-Sibe ...
.


Subspecies

*''A. a. allous'' (Geyer,
836 __NOTOC__ Year 836 ( DCCCXXXVI) was a leap year starting on Saturday (link will display the full calendar) of the Julian calendar. Events By place Abbasid Caliphate * Driven by tensions between his favoured Turkish guard and the populac ...
Alps, northern Europe *''A. a. vandalica'' Kaaber & Høegh-Guldberg, 1961 Jutland, Denmark *''A. a. rambringi'' Høegh-Guldberg, 1966 southern Scandinavia *''A. a. opheimi'' Høegh-Guldberg, 1966 southern Norway *''A. a. lyngensis'' Høegh-Guldberg, 1966 northern Scandinavia *''A. a. horkei'' Høegh-Guldberg, 1973 Öland & Gotland, Sweden *''A. a. ukrainica'' (Obraztsov, 1936) south-east Europe *''A. a. inhonora'' Jachontov, 1909 "Rossia ussiaentrali et orientali" *''A. a. sheljuzhkoi'' (Obraztsov, 1935) Caucasus Major *''A. a. turgaica'' (Obraztsov, 1935) south-west Siberia *''A. a. lepsinskana'' (Obraztsov, 1935) Dzhungarsky Alatau *''A. a. transalaica'' (Obratzov, 1935) Ghissar, Darvaz, Pamirs-Alai, Himalayas *''A. a. scytissa'' Nekrutenko, 1985 Tian-Shan *''A. a. sarmatis'' (Grum-Grshimailo, 1890) southern Urals *''A. a. strandi'' (Obraztsov, 1935) Altai – western Amur *''A. a. mandzhuriana'' (Obraztsov, 1935) eastern Amur, Ussuri *''A. a. hakutozana'' (Matsumura, 1927) North Korea *''A. a. sachalinensis'' (Matsumura, 1919) Sakhalin


Appearance, behaviour and distribution (Great Britain)

This species has confused British entomologists for years. Since its discovery in Britain it has been thought to be a form or a subspecies of the
brown argus ''Aricia agestis'', the brown argus, is a butterfly in the family Lycaenidae. It is found throughout the Palearctic realm, north to northern Jutland (Denmark) and east to Siberia and the Tian Shan. Subspecies *''A. a. agestis'' southern and ce ...
(''Aricia agestis'') and as well as a species in its own right. Since 1967 all brown arguses in the north of England and Scotland have been classified as this species. In the last few years genetic studies have shown that some of the colonies along the border of the two species ranges are still being wrongly classified and some colonies in the north of England are now thought to be ''A. agestis''. So far as is known their ranges do not overlap in the UK. The Scottish form is visually quite distinct from the brown argus. It usually has a small white dot in the centre of the upperside forewing and the black spots on the underside are missing leaving larger white spots on the light brown background. The northern English populations belong to a form called ''salmacis'' (called the Durham argus in English consulted 21 January 2014.) and are very similar to the brown argus including the presence of black spots on the under-wing (see photo) hence all the confusion. In Europe where it is known as the mountain argus, it is widespread in Scandinavia and mountainous regions of central, southern and eastern Europe. The species is considered locally rare in Britain, and the UK has established a detailed Biodiversity Action Plan to conserve this species along with a small number of other butterfly species.


Life cycle and food plants

:''Note that information on this species applies to Great Britain and some details may not be consistent with the species in other parts of its range.'' Eggs are laid singly on the upperside of the food plant leaves. As far is known common rock-rose (''Helianthemum nummularium'') seems to be the sole food plant in Britain. The larvae
hibernate Hibernation is a state of minimal activity and metabolic depression undergone by some animal species. Hibernation is a seasonal heterothermy characterized by low body-temperature, slow breathing and heart-rate, and low metabolic rate. It most ...
while still quite small and continue to feed and grow the following spring. Like the brown argus, it is attractive to ants and often attended by them. Pupation takes place at ground level in late May and butterflies are on the wing from mid-June to mid-July.


See also

*
List of butterflies of Great Britain This is a list of butterflies of Great Britain, including extinct, naturalised species and those of dubious origin. The list comprises butterfly species listed in ''The Moths and Butterflies of Great Britain and Ireland'' by Emmet ''et al.'' and '' ...
*
Fowlsheugh Fowlsheugh is a coastal nature reserve in Kincardineshire, northeast Scotland, known for its cliff formations and habitat supporting prolific seabird nesting colonies. Designated as a Site of Special Scientific Interest (SSSI) by Scottish Natur ...


References

{{Taxonbar, from=Q1549031 Aricia (butterfly) Butterflies of Europe Butterflies described in 1775