Argentine Espinal
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The Espinal (NT0801) is an
ecoregion An ecoregion (ecological region) or ecozone (ecological zone) is an ecologically and geographically defined area that is smaller than a bioregion, which in turn is smaller than a biogeographic realm. Ecoregions cover relatively large areas of l ...
of dry, thorny forest, savanna and steppe in Argentina. It has been extensively modified by large scale cattle ranching, but remnants of the original flora remain. It is threatened by the advance of the irrigation-based agricultural frontier.


Location

The espinal is a broad arc of arid land in the interior of Argentina to the north and west of
Buenos Aires Buenos Aires ( or ; ), officially the Autonomous City of Buenos Aires ( es, link=no, Ciudad Autónoma de Buenos Aires), is the capital and primate city of Argentina. The city is located on the western shore of the Río de la Plata, on South ...
, extending from the northern Uruguay border west through Santa Fe to Cordoba and south to the
Atlantic The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe an ...
coast south of
Bahía Blanca Bahía Blanca (; English: White Bay) is a city in the southwest of the provinces of Argentina, province of Buenos Aires Province, Buenos Aires, Argentina, by the Atlantic Ocean, and is the seat of government of the Bahía Blanca Partido. It had 3 ...
. It includes central
Santa Fe Province The Province of Santa Fe ( es, Provincia de Santa Fe, ) is a Provinces of Argentina, province of Argentina, located in the center-east of the country. Neighboring provinces are from the north clockwise Chaco Province, Chaco (divided by the 28th ...
, Córdoba Province and the northern part of
San Luis Province San Luis () is a province of Argentina located near the geographical center of the country (on the 32° South parallel). Neighboring provinces are, from the north clockwise, La Rioja, Córdoba, La Pampa, Mendoza and San Juan. History The ci ...
. It has an area of . The name "''espinal''" (thorny) refers to the thorny deciduous shrubland forest of the region between the flooded wetlands along the
Paraná River The Paraná River ( es, Río Paraná, links=no , pt, Rio Paraná, gn, Ysyry Parana) is a river in south-central South America, running through Brazil, Paraguay, and Argentina for some ."Parana River". Encyclopædia Britannica. Encyclopædia Br ...
to the east and the
Sierras de Córdoba The Sierras de Córdoba is a mountain range in central Argentina, located between the Pampas to the east and south and the Chaco to the north and east. Most of the range is located in Córdoba Province, except for the southwestern margin whic ...
to the west. The northern part of the ecoregion adjoins the
Humid Pampas The Humid Pampas ( es, Pampa Húmeda) is an extensive ecoregion of flat, fertile grassland of loessic origin in Argentina. It has a precipitation average of 900 mm per year, in contrast with the Dry Pampas to the west, which average less than ...
around Buenos Aires to the south, the
Uruguayan savanna The Uruguayan savanna, also known as the Brazilian-Uruguayan savanna, is a subtropical grassland and savanna ecoregion which includes all of Uruguay and southernmost Brazil, along with portions of Argentina along the Uruguay River, In Brazil, th ...
to the east, and the
Humid Chaco The Humid Chaco (Spanish: ''Chaco Húmedo'' or ''Chaco Oriental'') is tropical grasslands, savannas, and shrublands ecoregion in South America. It lies in the basin of the Paraná River, covering portions of central Paraguay and northern Argenti ...
and
Dry Chaco The Gran Chaco or Dry Chaco is a sparsely populated, hot and semiarid lowland natural region of the Río de la Plata basin, divided among eastern Bolivia, western Paraguay, northern Argentina, and a portion of the Brazilian states of Mato Gr ...
to the north and northwest. A belt of
Paraná flooded savanna The Paraná flooded savanna (NT0908) is an ecoregion that borders the southern Paraná River in Argentina. It has largely been converted to agriculture or occupied by urban development, but scattered patches of the original habitat remain along the ...
runs from north to south through the ecoregion along the Paraná River. The southern part of the ecoregion lies between the
Low Monte The Argentine Monte (NT0802), or Low Monte, is an ecoregion of dry thorn scrub and grasslands in Argentina. It is one of the driest regions in the country. Human settlements are mainly near water supplies such as rivers or oases. Deforestation and ...
ecoregion to the west and the Humid Pampas to the east.


Physical

The region mostly covers flat plains with
loess Loess (, ; from german: Löss ) is a clastic, predominantly silt-sized sediment that is formed by the accumulation of wind-blown dust. Ten percent of Earth's land area is covered by loess or similar deposits. Loess is a periglacial or aeolian ...
or sandy soils. There are wide variations in climate. The climate in the north is warm and wet, with rains in the summer. The
Köppen climate classification The Köppen climate classification is one of the most widely used climate classification systems. It was first published by German-Russian climatologist Wladimir Köppen (1846–1940) in 1884, with several later modifications by Köppen, notabl ...
is "Cfa": warm temperate, fully humid, hot summer. At a sample location at coordinates the mean temperature ranges from in July to in January. The yearly average mean temperature is about . Yearly total precipitation is about . Monthly precipitation ranges from in June to in March.


Ecology

The espinal is in the
Neotropical realm The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In bioge ...
, in the
temperate grasslands, savannas, and shrublands Temperate grasslands, savannas, and shrublands is a terrestrial biome defined by the World Wide Fund for Nature. The predominant vegetation in this biome consists of grass and/or shrubs. The climate is temperate and ranges from Semi-arid climate ...
biome. The dry ecoregion was once home to many species of birds, mammals and plants. The introduction of cattle in the 17th century had a huge impact, and the original habitat is now found only in isolated patches.


Flora

Vegetation includes deciduous
xerophytic A xerophyte (from Greek ξηρός ''xeros'' 'dry' + φυτόν ''phuton'' 'plant') is a species of plant that has adaptations to survive in an environment with little liquid water, such as a desert such as the Sahara or places in the Alps or th ...
forests, palm groves, grassy savannahs, grassy steppes and bushy steppes. Plants of the
Prosopis ''Prosopis'' is a genus of flowering plants in the family Fabaceae. It contains around 45 species of spiny trees and shrubs found in subtropical and tropical regions of the Americas, Africa, Western Asia, and South Asia. They often thrive in ari ...
and
Acacia ''Acacia'', commonly known as the wattles or acacias, is a large genus of shrubs and trees in the subfamily Mimosoideae of the pea family Fabaceae. Initially, it comprised a group of plant species native to Africa and Australasia. The genus na ...
genera of the
Mimosoideae The Mimosoideae are a traditional subfamily of trees, herbs, lianas, and shrubs in the pea family (Fabaceae) that mostly grow in tropical and subtropical climates. They are typically characterized by having radially symmetric flowers, with petals ...
family are common, add the mature forest areas are dominated by trees of the Prosopis genus. Other plants belong to genera such as
Jodina ''Jodina'' is a genus of flowering plants belonging to the family Santalaceae The Santalaceae, sandalwoods, are a widely distributed family of flowering plants (including small trees, shrubs, perennial herbs, and epiphytic climbersHewson & Ge ...
(
Santalaceae The Santalaceae, sandalwoods, are a widely distributed family of flowering plants (including small trees, shrubs, perennial herbs, and epiphytic climbersHewson & George t al.br>''Santalaceae'' taxonomy, 1984, pp. 191-194.) which, like other membe ...
family),
Celtis ''Celtis'' is a genus of about 60–70 species of deciduous trees, commonly known as hackberries or nettle trees, widespread in warm temperate regions of the Northern Hemisphere. The genus is part of the extended hemp family (Cannabaceae). De ...
,
Schinus ''Schinus'' is a genus of flowering trees and tall shrubs in the sumac family, Anacardiaceae. Members of the genus are commonly known as pepper trees. The Peruvian pepper tree (''Schinus molle'') is the source of the spice known as pink pepperco ...
, Geoffroea and Atamisquea (
Capparaceae The Capparaceae (or Capparidaceae), commonly known as the caper family, are a family of plants in the order Brassicales. As currently circumscribed, the family contains 33 genera and about 700 species. The largest genera are '' Capparis'' (about ...
family). In the ''carob'' subregion in the north of the espinal the main tree species include ''
Prosopis nigra ''Prosopis nigra'' (synonym ''Prosopis algarrobilla'') is a South American leguminous tree species that inhabits the Gran Chaco ecoregion (in particular, the transition zone between the Wet Chaco and the Southern Chaco), in Argentina and Paragua ...
'', ''
Prosopis alba ''Prosopis alba'' is a South American tree species that grows in central Argentina, the Gran Chaco ecoregion, and part of the Mesopotamia, Argentina, Argentine Mesopotamia, as well as Bolivia, Paraguay, and Peru. It is known as ''algarrobo blanc ...
'', ''
Celtis tala ''Celtis tala'' (or ''Celtis ehrenbergiana''), known as tala, is a medium size deciduous tree, native to tropical and subtropical South America. With small to medium-sized spines, its one of the main components of the Gran Chaco prairies and cer ...
'', ''
Geoffroea decorticans ''Geoffroea decorticans'', the chañar, kumbaru, or Chilean palo verde (green wood), is a small deciduous tree, up to 8 meters (25 ft) tall that inhabits most arid forests (montes or espinales) of southern South America. The chañar is cold ...
'', ''
Vachellia caven ''Vachellia caven'' (Roman cassie, , aromo criollo, caven, , , espinillo, espinillo de baado, espino, espino maulino) is an ornamental tree in the family Fabaceae. ''Vachellia caven'' is native to Argentina, Bolivia, Chile, Paraguay, and Urugua ...
'', ''
Jodina rhombifolia ''Jodina rhombifolia'' (the '' quebracho flojo'' (the loose quebracho), ''quebracho flajo'', ''sombra de toro'' or ''quebrachillo'') is a tree species in the family Santalaceae. References External links Sombra de toro image on flickr Santa ...
'', '' Scutia buxifolia'' and '' Schinus longifolia. Isolated specimens or groups of the '' Trithrinax campestris'' palm are still found in some areas. The "caldén" subregion in the south of the espinal has xerophytic forests with many ''
Prosopis caldenia ''Prosopis caldenia'', commonly known as the caldén, is a species of flowering tree in the family Fabaceae The Fabaceae or Leguminosae,halophilic The halophiles, named after the Greek word for "salt-loving", are extremophiles that thrive in high salt concentrations. While most halophiles are classified into the domain Archaea, there are also bacterial halophiles and some eukaryotic species, ...
steppes.


Fauna

There are three species of endemic birds. The
Mar Chiquita Mar Chiquita is a coastal lagoon in the southeast province of Buenos Aires in eastern Argentina. It is located by the Atlantic coast, 30 km north of Mar del Plata. The area is a natural reserve where a number of animal species live around ...
lake and the Dulce River swamps are designated a
Ramsar wetland The Ramsar Convention on Wetlands of International Importance Especially as Waterfowl Habitat is an international treaty for the conservation and sustainable use of Ramsar sites (wetlands). It is also known as the Convention on Wetlands. It i ...
that is a key site for endangered neotropical birds. The wetland supports a rich and diverse fauna with 138 known species of birds, including large breeding groups of
Chilean flamingo The Chilean flamingo (''Phoenicopterus chilensis'') is a species of large flamingo at closely related to the American flamingo and the greater flamingo, with which it was sometimes considered conspecific. The species is listed as near threatened ...
(''Phoenicopterus chilensis''). Native species include
guanaco The guanaco (; ''Lama guanicoe'') is a camelid native to South America, closely related to the llama. Guanacos are one of two wild South American camelids, the other being the vicuña, which lives at higher elevations. Etymology The guanaco g ...
s,
greater rhea The greater rhea (''Rhea americana'') is a species of flightless bird native to eastern South America. Other names for the greater rhea include the grey, common, or American rhea; ema (Portuguese); or ñandú (Guaraní and Spanish). One of two sp ...
s, and
giant anteater The giant anteater (''Myrmecophaga tridactyla'') is an insectivorous mammal native to Central and South America. It is one of four living species of anteaters, of which it is the largest member. The only extant member of the genus ''Myrmecopha ...
s. Endangered amphibians include blunt-headed salamander (''Ambystoma amblycephalum''). Endangered birds include yellow cardinal (''Gubernatrix cristata''),
Chaco eagle The Chaco eagle (''Buteogallus coronatus'') or crowned solitary eagle, is an endangered bird of prey from eastern and central South America. Typically it is known simply as the crowned eagle which leads to potential confusion with the African ''S ...
(''Buteogallus coronatus'') and
Eskimo curlew The Eskimo curlew (''Numenius borealis''), also known as northern curlew, is a species of curlew in the family Scolopacidae. It was one of the most numerous shorebirds in the tundra of western Arctic Canada and Alaska. Thousands of birds were th ...
(''Numenius borealis'').


Status

The
World Wildlife Fund The World Wide Fund for Nature Inc. (WWF) is an international non-governmental organization founded in 1961 that works in the field of wilderness preservation and the reduction of human impact on the environment. It was formerly named the Wo ...
gives the espinal ecoregion the status of "Critical/Endangered". Much of the ecoregion has been used for agriculture, and the trees of the sparse dry forests have been used for firewood, fence posts, flooring and furniture. Irrigation systems are expanding the agricultural frontier into area that were previously not affected. The Dulce River floodplain and Mar Chiquita lake are threatened by increasing use of the water upstream. In parts of the ecoregion, woody plant encroachment has been observed.


Protected areas

1.74% of the ecoregion is in protected areas. They include the
Lihué Calel National Park Lihué Calel National Park ( es, Parque Nacional Lihué Calel) ( Mapudungun Lihue = "life" and Calel "mountainous area", "mountains of life") is a national park in Argentina Argentina (), officially the Argentine Republic ( es, link=no ...
,
La Reforma University Reserve LA most frequently refers to Los Angeles, the second largest city in the United States. La, LA, or L.A. may also refer to: Arts and entertainment Music * La (musical note), or A, the sixth note * "L.A.", a song by Elliott Smith on ''Figur ...
, Chacharramendi Provincial Reserve and Luro Provincial Reserve. Other units are the
El Palmar National Park El Palmar National Park (in Spanish, ''Parque Nacional El Palmar'') is one of Argentina's national parks, located on the center-east of the province of Entre Ríos, midway between the cities of Colón (54 km) and Concordia (60 km). I ...
and the Parque Atlántico Mar Chiquita.


Notes


References


Sources

* * * {{DEFAULTSORT:Espinal Neotropical ecoregions Ecoregions of Argentina Temperate grasslands, savannas, and shrublands