Arctogeophilus Glacialis
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''Arctogeophilus glacialis'' is a species of
soil centipede Centipedes (from New Latin , "hundred", and Latin , "foot") are predatory arthropods belonging to the class Chilopoda (Ancient Greek , ''kheilos'', lip, and New Latin suffix , "foot", describing the forcipules) of the subphylum Myriapoda, an a ...
in the family Geophildae. This
centipede Centipedes (from New Latin , "hundred", and Latin , " foot") are predatory arthropods belonging to the class Chilopoda (Ancient Greek , ''kheilos'', lip, and New Latin suffix , "foot", describing the forcipules) of the subphylum Myriapoda, an ...
is found in Russia, Alaska, and Canada. This species was first described in 1909 by the Austrian
myriapodologist Myriapodology is the scientific study of myriapods which includes centipedes and millipedes. The field of myriapodology can also cover other myriapods such as pauropods and symphylans. Those who study myriapods are myriapodologists. Societies * In ...
Carl Attems Carl August Graf Attems-Petzenstein (13 October 1868 in Graz, Austria – 19 April 1952 in Vienna) was an Austrian myriapodologist and invertebrate zoologist. He published 138 scientific papers, most of them dealing with his specialist field ...
as the type species for the taxon '' Arctogeophilus'', which was originally described as a
subgenus In biology, a subgenus (plural: subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between t ...
of the genus ''
Geophilus ''Geophilus '' is a large, heterogeneous genus of soil centipedes in the family Geophilidae largely considered to be synonymous with '' Brachygeophilus''. It is a mostly holarctic genus characterized by a claw-shaped ultimate pretarsus, anteri ...
'' but was elevated to the status of genus in 1910 by the French zoologist Henri Ribaut.


Discovery, taxonomy, and distribution

Attems based the original description of this species on several
type specimens In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to anchor or centralizes t ...
, including three males and four females. These
syntypes In biological nomenclature, a syntype is any one of two or more biological types that is listed in a description of a taxon where no holotype was designated. Precise definitions of this and related terms for types have been established as part of ...
were found in 1879 at
Port Clarence Port Clarence is a small village now within the borough of Stockton-on-Tees and ceremonial county of County Durham, England. It is situated on the north bank of the River Tees, and hosts the northern end of the Middlesbrough Transporter Bridge. ...
in Alaska and at two sites on the Chukotka peninsula (the village of Nunyamo and
Penkigney Bay Penkigney Bay ( rus, Пенкегней; Chukchi: Пэнкэнэй) is a bay of the Bering Sea on the eastern coast of the Chukotka Peninsula, Russian Federation. Administratively the bay is part of the Providensky District of Chukotka. Geogra ...
) on the eastern coast of Siberia. These specimens were collected during the first successful navigation of the
Northeast Passage The Northeast Passage (abbreviated as NEP) is the shipping route between the Atlantic and Pacific Oceans, along the Arctic coasts of Norway and Russia. The western route through the islands of Canada is accordingly called the Northwest Passage (N ...
, completed by the Vega expedition led by the Nordic explorer
Adolf Erik Nordenskjöld Adolf (also spelt Adolph or Adolphe, Adolfo and when Latinised Adolphus) is a given name used in German-speaking countries, Scandinavia, the Netherlands and Flanders, France, Italy, Spain, Portugal, Latin America and to a lesser extent in vari ...
aboard the
SS Vega A number of steamships have carried the name ''Vega'', including *, Adolf Erik Nordenskiöld sailed in 1878 in ''Vega'' from Gothenburg along the coast of Siberia to Yokohama on his discovery of the Northeast Passage. *SS ''Vega'', launched in 18 ...
. These specimens and other
type material In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to anchor or centralizes the ...
in the form of slides are deposited in the Natural History Museum in Vienna. In 1919, the American
biologist A biologist is a scientist who conducts research in biology. Biologists are interested in studying life on Earth, whether it is an individual cell, a multicellular organism, or a community of interacting populations. They usually specialize in ...
Ralph V. Chamberlain described ''Cryophilus alaskanus'' as a new species based on two specimens collected in 1916 from the tundra near Nome in Alaska. In 1946, Chamberlain reported the collection of another specimen in 1945 in College near
Fairbanks Fairbanks is a home rule city and the borough seat of the Fairbanks North Star Borough in the U.S. state of Alaska. Fairbanks is the largest city in the Interior region of Alaska and the second largest in the state. The 2020 Census put the po ...
in Alaska, but he also deemed ''C. alaskanus'' to be a
junior synonym The Botanical and Zoological Codes of nomenclature treat the concept of synonymy differently. * In botanical nomenclature, a synonym is a scientific name that applies to a taxon that (now) goes by a different scientific name. For example, Linna ...
of ''A. glacialis''. Authorities now consider these centipedes to be the same species. In 2022, David W. Langor and Stephen D. Langor reported the discovery of more than 20 specimens in Canada. These centipedes were recently identified as specimens of ''A. glacialis'' by the American myriapodologist Ralph E. Crabill, Jr. These specimens were collected at Reindeer Station in the
Northwest Territories The Northwest Territories (abbreviated ''NT'' or ''NWT''; french: Territoires du Nord-Ouest, formerly ''North-Western Territory'' and ''North-West Territories'' and namely shortened as ''Northwest Territory'') is a federal territory of Canada. ...
by the Canadian zoologist John R. Vockeroth in 1948 and represent the first record of this species in Canada.


Description

This species features 39 pairs of legs in each sex and can reach 20 mm in length and 1.2 mm in breadth. This centipede has a yellow or reddish yellow body with the head more reddish brown than the body. The first
maxillae The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The t ...
feature relatively short lappets projecting from the lateral margins. The
forcipules Forcipules are the modified, pincer-like, front legs of centipedes that are used to inject venom into prey. They are the only known examples of front legs acting as venom injectors. Nomenclature Forcipules go by a variety of names in both scien ...
feature denticles on all articles, with the denticles on the second and third articles only slightly shorter than those on the first and ultimate articles. The sterna lack ventral fields of pores, and each of the
ultimate legs Ultimate legs are a pair of modified rear legs unique to centipedes. Although they do not aid in locomotion, ultimate legs are used for a variety of uses, and their morphology varies accordingly. Sexual dimorphism is frequently present. Usage ...
lacks a claw or pretarsus at the tip. This species shares many traits with other ''Arctogeophilus'' species. For example, centipedes in this genus usually lack ventral pore-fields and feature no pretarsus on the ultimate legs. Like other species in this genus, ''A. glacialis'' features an elongate head, with a cephalic plate that is noticeably longer than wide. Furthermore, the labrum in this genus features side pieces that nearly touch in the middle, making the intermediate part inconspicuous. In ''A. glacialis'', the intermediate part of the labrum is covered by the side pieces, which meet in the middle. Three other ''Arctogeophilus'' species found in Russia, ''A. attemsi'', '' A. macrocephalus'', and '' A. sachalinus'', are so similar to ''A. glacialis'' that some authors have suggested that these three may be junior synonyms of ''A. glacialis''. The species ''A. glacialis'' may be distinguished from these close relatives, however, by the lappets on the first maxillae, the denticles on the forcipular articles, and the absence of ventral pore-fields. Whereas the first maxillary lappets are short in ''A. glacialis'', they are long in the other three species. Furthermore, denticles are present on the second and third articles of the forcipule in ''A. glacialis'', whereas these denticles are absent in ''A. attemsi'' and ''A. sachalinus''. Moreover, ventral pore fields appear on some anterior segments in ''A. macrocephalus'' and ''A. sachalinus'' but are entirely absent in ''A. glacialis''.


References

{{Taxonbar, from=Q6505484 Geophilidae Taxa named by Carl Attems Animals described in 1909