Archilestes Grandis
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The great spreadwing (''Archilestes grandis'') is a
damselfly Damselflies are flying insects of the suborder Zygoptera in the order Odonata. They are similar to dragonflies, which constitute the other odonatan suborder, Anisoptera, but are smaller and have slimmer bodies. Most species fold the wings along ...
in the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Lestidae The Lestidae are a rather small family of cosmopolitan, large-sized, slender damselflies, known commonly as the spreadwings or spread-winged damselflies. Characteristics While most damselflies rest with their wings folded together, most member ...
. When great spreadwings are startled they often return to the same perch or a perch nearby.


Characteristics

The great spreadwing is one of the largest North American spreadwings, with a length of 2-2.4 inches and a wingspan of 3 inches. The
thorax The thorax or chest is a part of the anatomy of humans, mammals, and other tetrapod animals located between the neck and the abdomen. In insects, crustaceans, and the extinct trilobites, the thorax is one of the three main divisions of the cre ...
of the male is dull greenish bronze above it is a broad diagonal yellow stripe on sides. It is also the only species with a broad yellow racing stripe on the sides of thorax. The
abdomen The abdomen (colloquially called the belly, tummy, midriff, tucky or stomach) is the part of the body between the thorax (chest) and pelvis, in humans and in other vertebrates. The abdomen is the front part of the abdominal segment of the torso. ...
is dark with a blue-gray tip. Its eyes and face are blue. Females are similar to males but are more brown on the body. Her eyes are more of a paler blue than the male. The yellow stripe also occurs on the female great spreadwing. When females are laying eggs they may appear in a putty-color. It is much the same color as the withered leaves in which they lay eggs.


Distribution

The great spreadwing is found in western and southern North America. It is more common in western United States than in the eastern. The great spreadwing can also be found in Central America and northern South America. In the 1920s and later ''Archilestes grandis'' has recently expanded its range in North America to the east and north.


Habitat

The great spreadwing prefers slow small streams, often with
alder Alders are trees comprising the genus ''Alnus'' in the birch family Betulaceae. The genus comprises about 35 species of monoecious trees and shrubs, a few reaching a large size, distributed throughout the north temperate zone with a few sp ...
or
willow Willows, also called sallows and osiers, from the genus ''Salix'', comprise around 400 speciesMabberley, D.J. 1997. The Plant Book, Cambridge University Press #2: Cambridge. of typically deciduous trees and shrubs, found primarily on moist s ...
s, wetlands, ponds and temporary pools. It breeds readily in heavily vegetated water gardens with unpolluted water, as well as in natural water. Females will
oviposit The ovipositor is a tube-like organ used by some animals, especially insects, for the laying of eggs. In insects, an ovipositor consists of a maximum of three pairs of appendages. The details and morphology of the ovipositor vary, but typical ...
in vegetation such as water iris while clasped by males. The male great spreadwings hanging from vegetation either over or near water, usually fairly low, like under 2 feet. On an occasion the
nymph A nymph ( grc, νύμφη, nýmphē, el, script=Latn, nímfi, label=Modern Greek; , ) in ancient Greek folklore is a minor female nature deity. Different from Greek goddesses, nymphs are generally regarded as personifications of nature, are ty ...
s can be spotted in winter, basking in sunwarmed shallow water alongside tadpoles. They are recognizable by their Y tails which are actually their gills.


Flight season

This species is most active from April–December. The timing varies with temperature.


Life cycle

When great spreadwings are mating and after collection of sperm, the female cuts a slit in emergent vegetation and oviposits in it. Then nymphs develop underwater until they climb out, ready to emerge as adults. Like other Odonates, emergence usually occurs under the cover of darkness.


References

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Great Spreadwing - Archilestes grandis - BugGuide.Net
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Wisconsin Odonata Survey - Archilestes grandis
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Great Spreadwing - Archilestes grandis
{{Taxonbar, from=Q2206513 Lestidae Odonata of North America Insects of the United States Fauna of the Western United States