Arachnothera
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The spiderhunters are birds of the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''Arachnothera'', part of the
sunbird Sunbirds and spiderhunters make up the family Nectariniidae of passerine birds. They are small, slender passerines from the Old World, usually with downward-curved bills. Many are brightly coloured, often with iridescent feathers, particularly i ...
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Nectariniidae Sunbirds and spiderhunters make up the family Nectariniidae of passerine birds. They are small, slender passerines from the Old World, usually with downward-curved bills. Many are brightly coloured, often with iridescent feathers, particularly i ...
. The genus contains thirteen species found in the forests of south and southeastern Asia. They are large representatives of the sunbird family, with drab plumage and long strongly curved bills. They feed on both nectar and a range of small arthropods.


Distribution and habitat

Unlike the rest of the family, which is more widespread, the spiderhunters are confined to the Oriental zoogeographic region, occurring from
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
east to the
Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
and from the
Himalayas The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the planet's highest peaks, including the very highest, Mount Everest. Over 100 ...
south to
Java Java (; id, Jawa, ; jv, ꦗꦮ; su, ) is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea to the north. With a population of 151.6 million people, Java is the world's List ...
; they reach their greatest species diversity in the
Thai-Malay peninsula The Malay Peninsula ( Malay: ''Semenanjung Tanah Melayu'') is a peninsula in Mainland Southeast Asia. The landmass runs approximately north–south, and at its terminus, it is the southernmost point of the Asian continental mainland. The ar ...
,
Sumatra Sumatra is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the sixth-largest island in the world at 473,481 km2 (182,812 mi.2), not including adjacent i ...
and
Borneo Borneo (; id, Kalimantan) is the third-largest island in the world and the largest in Asia. At the geographic centre of Maritime Southeast Asia, in relation to major Indonesian islands, it is located north of Java, west of Sulawesi, and eas ...
. The spiderhunters are mostly forest birds, occupying a wide range of forest types including true rainforest,
dipterocarp Dipterocarpaceae is a family of 16 genera and about 695 known species of mainly tropical lowland rainforest trees. The family name, from the type genus ''Dipterocarpus'', is derived from Greek (''di'' = two, ''pteron'' = wing and ''karpos'' = fru ...
forest, swamp forest, bamboo forest, secondary forest, forest edge and other highly degraded forest. In addition several species occur in human-created habitat such as gardens or plantations. Most species are lowland species, but the Whitehead's spiderhunter is more of a montane species.


Description

The spiderhunters are amongst the largest of their family, measuring between 13 and 22 cm in length. The
spectacled spiderhunter The spectacled spiderhunter (''Arachnothera flavigaster'') is a species of bird in the family Nectariniidae. It is found in Brunei, Indonesia, Malaysia, Singapore, Thailand, and Vietnam. Its natural habitats are subtropical or tropical moist lowl ...
is the heaviest of the sunbirds, weighing as much as 49 g. The bills of the spiderhunters are long, at least twice the length of the head, and are decurved and stout. The tongue forms a complete tube for most of its length. The
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
is much drabber than the other brightly coloured sunbirds, is the same for both sexes, and contains no
iridescence Iridescence (also known as goniochromism) is the phenomenon of certain surfaces that appear to gradually change color as the angle of view or the angle of illumination changes. Examples of iridescence include soap bubbles, feathers, butterfl ...
. The uppersides of most species are olive-green and the undersides dull white or yellow - in half the species the undersides are streaked. The most atypical spiderhunter plumage is that of the Whitehead's spiderhunter, which is dark brown with white streaking on the undersides and back and a yellow rump. The calls of the spiderhunters are very simple, typically a metallic ''chip'' which is repeated multiple times to form the song. The song of the little spiderhunter is described as an "incessant squeaky whistle".


Diet and feeding

The spiderhunters are
omnivorous An omnivore () is an animal that has the ability to eat and survive on both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize the nutri ...
. As their name suggests they will eat spiders, and are capable of extracting spiders from the centre of their webs, a tricky task. They also consume a wide range of other small arthropod prey including crickets, caterpillars, butterflies, ants and other insects. In addition to animal prey they also consume
nectar Nectar is a sugar-rich liquid produced by plants in glands called nectaries or nectarines, either within the flowers with which it attracts pollinating animals, or by extrafloral nectaries, which provide a nutrient source to animal mutualists ...
from flowers. Their tubular tongue is pushed against the top of the upper mandible of the bill, then pulled in and out, creating a pressure difference which allows the nectar to be sucked into the mouth. Spiderhunters may be important
pollinator A pollinator is an animal that moves pollen from the male anther of a flower to the female stigma of a flower. This helps to bring about fertilization of the ovules in the flower by the male gametes from the pollen grains. Insects are the maj ...
s of some species of plants, and species pollinated by spiderhunters have long tubular flowers. They may also participate in
nectar robbing Nectar robbing is a foraging behavior utilized by some organisms that feed on floral nectar, carried out by feeding from holes bitten in flowers, rather than by entering through the flowers' natural openings. "Nectar robbers" usually feed in this ...
, which consists of inserting the bill into the side of the flower to extract nectar without being dabbed with pollen. Little is known about their territorial behaviour, but there have been reports of some species defending territories. The spiderhunters are less gregarious than other sunbirds, and are usually seen alone or as part of a pair.


Breeding

The spiderhunters are thought to be
monogamous Monogamy ( ) is a form of Dyad (sociology), dyadic Intimate relationship, relationship in which an individual has only one Significant other, partner during their lifetime. Alternately, only one partner at any one time (Monogamy#Serial monogamy, ...
like most of the rest of the family. They vary from the other sunbirds in the nature of their
nest A nest is a structure built for certain animals to hold eggs or young. Although nests are most closely associated with birds, members of all classes of vertebrates and some invertebrates construct nests. They may be composed of organic materia ...
, which is suspended underneath a large leaf, usually a banana leaf but sometimes a palm frond or even a branch. The style of nest can be quite variable, a simple cup in the case of the
streaked spiderhunter The streaked spiderhunter (''Arachnothera magna'') is a species of bird in the family Nectariniidae. Description It is similar in size to a sparrow. It is olive yellow in color, with multiple darker streaks. It has a long, curved black beak and ...
, an elongated tube for the
yellow-eared spiderhunter The yellow-eared spiderhunter (''Arachnothera chrysogenys'') is a species of bird in the family Nectariniidae. It is found in Brunei, Indonesia, Malaysia, Myanmar, Singapore, Thailand, and Vietnam. Its natural habitats are subtropical or tropical ...
and a bottle shape for the long-billed spiderhunter. The nests are suspended from the leaf by using spider web or by pushing plant fibres through the leaf. The nests are made of grasses and leaves and lined with soft materials. Nest construction, as is typical of the family, is carried out by the female alone. Unlike the other sunbirds, however, both the male and female incubate the eggs. The spiderhunters lay two or three eggs. Their nests are sometimes
parasitised Parasitism is a close relationship between species, where one organism, the parasite, lives on or inside another organism, the host, causing it some harm, and is adapted structurally to this way of life. The entomologist E. O. Wilson has ...
by
cuckoo Cuckoos are birds in the Cuculidae family, the sole taxon in the order Cuculiformes . The cuckoo family includes the common or European cuckoo, roadrunners, koels, malkohas, couas, coucals and anis. The coucals and anis are sometimes separ ...
s.


Species

The genus contains 13 species:


References


External links


Spiderhunter videos
on the Internet Bird Collection {{Taxonbar, from=Q774692 Arachnothera   Taxa named by Coenraad Jacob Temminck