Arachnomorpha is a proposed subdivision or

clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...

Arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arth ...

a, comprising the group formed by the

trilobites Trilobites (; meaning "three lobes") are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the At ...

and their close relatives (

Artiopoda The Artiopoda is a grouping of extinct arthropods that includes trilobites and their close relatives. It was erected by Hou and Bergström in 1997 to encompass a wide diversity of arthropods that would traditionally have been assigned to the Trilo ...

Megacheira Megacheira ("great hands") is an extinct class of predatory arthropods that possessed a pair of great appendages, hence the class name as well as the common name "great appendage arthropods". Their taxonomic position is controversial, with stud ...

(which may be

paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...

) and

chelicerates The subphylum Chelicerata (from New Latin, , ) constitutes one of the major subdivisions of the phylum Arthropoda. It contains the sea spiders, horseshoe crabs, and arachnids (including harvestmen, scorpions, spiders, solifuges, ticks, and mit ...

. Under this proposed classification scheme, Arachnomorpha is considered the sister group to

Mandibulata Mandibulata, termed "mandibulates", is a clade of arthropods that comprises the extant subphyla Myriapoda (millipedes and others), Crustacea and Hexapoda (insects and others). Mandibulata is currently believed to be the sister group of the clade ...

(including

insects Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (head, thorax and abdomen), three pairs of j ...

crustaceans Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...

and

myriapods Myriapods () are the members of subphylum Myriapoda, containing arthropods such as millipedes and centipedes. The group contains about 13,000 species, all of them terrestrial. The fossil record of myriapods reaches back into the late Silurian, a ...

). The arachnomorph concept has been challenged by suggestions that the artiopods are more closely related to mandibulates, forming the clade Antennulata instead. There is no consensus as to assigning Arachnomorpha a formal Linnean rank.

Classification

Arachnomorpha Lameere, 1890 Arachnata Paulus, 1979, = Palaeopoda Packard, 1903*†

Trilobita Trilobites (; meaning "three lobes") are extinction, extinct marine arthropods that form the class (biology), class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil reco ...

Walch, 1771 *†

Hou & Bergstrom, 1997 *†

Strabopida Strabopidae is the only family of the order Strabopida, a group of arachnomorph arthropods. The order Strabopida contains three valid genera and one doubtful, ''Khankaspis'', '' Paleomerus'', '' Parapaleomerus'' and '' Strabops'', all assigned in ...

Hou & Bergström, 1997 *† Aglaspida Walcott, 1911 *† Cheloniellida Broili, 1932 *

Chelicerata The subphylum Chelicerata (from New Latin, , ) constitutes one of the major subdivisions of the phylum Arthropoda. It contains the sea spiders, horseshoe crabs, and arachnids (including Opiliones, harvestmen, scorpions, spiders, Solifugae, solif ...

Heymons 1901 **Family † Sanctacarididae Legg & Pates, 2017 **Class

Pycnogonida Sea spiders are marine arthropods of the order Pantopoda ( ‘all feet’), belonging to the class Pycnogonida, hence they are also called pycnogonids (; named after ''Pycnogonum'', the type genus; with the suffix '). They are cosmopolitan, fou ...

Latreille, 1810 **Clade

Euchelicerata The subphylum Chelicerata (from New Latin, , ) constitutes one of the major subdivisions of the phylum Arthropoda. It contains the sea spiders, horseshoe crabs, and arachnids (including harvestmen, scorpions, spiders, solifuges, ticks, and mite ...

Weygoldt & Paulus, 1979 ***Genus †

Offacolidae ''Offacolus'' is an extinct genus of euchelicerate, a group of chelicerate arthropods. Its only species, ''O. kingi'', has been found in deposits from the Silurian period (Homerian epoch) in the Wenlock Series Lagerstätte of Herefordshire, Eng ...

Sutton ''et al''., 2002 ***Clade

Prosomapoda Prosomapoda is a clade of euchelicerates including the groups Xiphosura (horseshoe crabs) and Planaterga (a group comprising bunodids, pseudoniscids, chasmataspidids, eurypterids and arachnids), as well as several basal synziphosurid genera ...

Lamsdell, 2013

Phylogeny

Using fossil data, Bergström & Hou (2003) gave an outline of arthropod relationships emphasizing trilobitomorphs (a group that includes trilobites and trilobite-like animals).

References

Arthropod taxonomy Taxa named by Auguste Lameere {{arthropod-stub