Arachniodes × Minamitanii
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''Arachniodes'' is a
fern A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except t ...
genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the
Pteridophyte Phylogeny Group The Pteridophyte Phylogeny Group, or PPG, is an informal international group of systematic botanists who collaborate to establish a consensus on the classification of pteridophytes (lycophytes and ferns) that reflects knowledge about plant relation ...
classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns". The genus ''Arachniodes'' was first published by Carl Ludwig von Blume in 1828, with the single Indonesian species ''Arachniodes aspidioides''. The genus was not widely recognized until Mary Douglas Tindale transferred the two species (''Byrsopteris amabilis'' and ''Byrsopteris aristata'') into it in 1961.Tindale, Mary Douglas; ''Contributions from the New South Wales National Herbarium'' 3(1): 90. 1961.


Species

The
Pteridophyte Phylogeny Group The Pteridophyte Phylogeny Group, or PPG, is an informal international group of systematic botanists who collaborate to establish a consensus on the classification of pteridophytes (lycophytes and ferns) that reflects knowledge about plant relation ...
classification of 2016 (PPG I) considers ''Arachniodes'' to have 60 species. , the ''Checklist of Ferns and Lycophytes of the World'' accepted 78 species: *'' Arachniodes ailaoshanensis'' Ching *'' Arachniodes amabilis'' (Blume) Tindale *''
Arachniodes amoena ''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns". The genus ...
'' (Ching) Ching *'' Arachniodes aristata'' (G.Forst.) Tindale *'' Arachniodes assamica'' (Kuhn) Ohwi *'' Arachniodes bella'' (C.Chr.) Ching *'' Arachniodes blinii'' (H.Lév.) Nakaike *'' Arachniodes carvifolia'' (Kunze) Ching *''
Arachniodes caudata ''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns". The genus ...
'' Ching *'' Arachniodes cavaleriei'' (Christ) Ohwi *''
Arachniodes chinensis ''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns". The genus ' ...
'' (Rosenst.) Ching *''
Arachniodes coniifolia ''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns". The genus ' ...
'' (T.Moore) Ching *'' Arachniodes cornucervi'' (D.Don) Fraser-Jenk. *'' Arachniodes daklakensis'' Li Bing Zhang, N.T.Lu & X.M.Zhou *'' Arachniodes davalliaeformis'' (Christ) Nakaike *''
Arachniodes denticulata ''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns". The genus ' ...
'' (Sw.) Ching *''
Arachniodes fengii ''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns". The genus ' ...
'' Ching *''
Arachniodes festina ''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns". The genus ' ...
'' (Hance) Ching *''
Arachniodes formosa ''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns". The genus ' ...
'' (Fée) Ching *''
Arachniodes formosissima ''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns". The genus ' ...
'' (Goldm.) Nakaike *''
Arachniodes gigantea ''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns". The genus ' ...
'' Ching *''
Arachniodes globisora ''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns". The genus ' ...
'' (Hayata) Ching *''
Arachniodes grossa ''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns". The genus ' ...
'' (Tardieu & C.Chr.) Ching *''
Arachniodes hainanensis ''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns". The genus ' ...
'' (Ching) Ching *''
Arachniodes haniffii ''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns". The genus ' ...
'' (Holttum) Ching *''
Arachniodes hehaii ''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns". The genus ' ...
'' Li Bing Zhang, N.T.Lu & X.F.Gao *''
Arachniodes hekiana ''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns". The genus ...
'' Kurata *''
Arachniodes henryi ''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns". The genus ...
'' (Christ) Ching *'' Arachniodes hiugana'' Sa. Kurata *''
Arachniodes hunanensis ''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns". The genus ' ...
'' Ching *'' Arachniodes insularis'' W.H.Wagner *'' Arachniodes japonica'' (Kurata) Nakaike *'' Arachniodes jinpingensis'' Y.T.Hsieh *'' Arachniodes leucostegioides'' (C.Chr.) Ching *'' Arachniodes longicaudata'' Li Bing Zhang, N.T.Lu & Liang Zhang *'' Arachniodes longipinna'' Ching *'' Arachniodes microlepioides'' (C.Chr.) comb. ined. *''
Arachniodes miqueliana ''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns". The genus ...
'' (Maxim. ex Franch. & Sav.) Ohwi *'' Arachniodes miyakei'' (H.Itô) Shimura ex Nakaike *'' Arachniodes mutica'' (Franch. & Sav.) Ohwi *'' Arachniodes neopodophylla'' (Ching) Nakaike *'' Arachniodes nigrospinosa'' (Ching) Ching *'' Arachniodes nipponica'' (Rosenst.) Ohwi *'' Arachniodes oosorae'' H.Itô *'' Arachniodes palmipes'' (Kunze) Fraser-Jenk. *'' Arachniodes pseudoaristata'' (Tagawa) Ohwi *'' Arachniodes pseudoassamica'' Ching *'' Arachniodes pseudorepens'' Nakaike *'' Arachniodes puncticulata'' (Alderw.) Ching *'' Arachniodes quadripinnata'' (Hayata) Seriz. *'' Arachniodes quangnamensis'' Li Bing Zhang, N.T.Lu & Liang Zhang *'' Arachniodes repens'' Kurata *''
Arachniodes rhomboidea ''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns". The genus ...
'' (Schott) Ching *''
Arachniodes rigidissima ''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns". The genus ...
'' (Hook.) Ching *'' Arachniodes sarasiniorum'' (Christ) Nakaike *''
Arachniodes simplicior ''Arachniodes simplicior'', the simpler East Indian hollyfern, East Indian holly fern, or variegated shield fern, is a species of fern in the family Dryopteridaceae. Distribution China (Anhui, Chongqing, Fujian, SE-Gansu, Guangxi, Guizhou, Henan ...
'' (Makino) Ohwi *''
Arachniodes simulans ''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns". The genus ...
'' (Ching) Ching *'' Arachniodes sinomiqueliana'' (Ching) Ohwi *'' Arachniodes sinorhomboidea'' Ching *'' Arachniodes sledgei'' Fraser-Jenk. *''
Arachniodes spectabilis ''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns". The genus ...
'' (Ching) Ching *''
Arachniodes squamulosa ''Arachniodes squamulosa'' is a species of fern in the family Dryopteridaceae. It is endemic to Ecuador. Its natural habitats are subtropical or tropical moist lowland forests and subtropical or tropical moist montane forests. It is threatened b ...
'' R.C.Moran & B. Øllg. *'' Arachniodes standishii'' (T.Moore) Ohwi *'' Arachniodes subamabilis'' Kurata *'' Arachniodes superba'' Fraser-Jenk. *'' Arachniodes tomitae'' Sa. Kurata *''
Arachniodes tonkinensis ''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns". The genus ...
'' (Ching) Ching *'' Arachniodes tripinnata'' (Goldm.) Sledge *'' Arachniodes tsiangiana'' (Ching) Nakaike *''
Arachniodes webbiana ''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns". The genus ...
'' (A.Braun) Schelpe *'' Arachniodes wulingshanensis'' S.F.Wu *'' Arachniodes yakusimensis'' (H.Itô) Nakaike *'' Arachniodes ziyunshanensis'' Y.T.Xie Some hybrids were also accepted: * ''Arachniodes'' × ''azuminoensis'' Fujiw., Y.Matsuda, Yu. Abe & Otsuka * ''Arachniodes'' × ''chibaensis'' Yashiro * ''Arachniodes'' × ''clivorum'' Kurata * ''Arachniodes'' × ''ikeminensis'' Seriz. * ''Arachniodes'' × ''kenzo-satakei'' (Kurata) Kurata * ''Arachniodes'' × ''kurosawae'' Shimura & Kurata * ''Arachniodes'' × ''masakii'' Kurata * ''Arachniodes'' × ''minamitanii'' Sa. Kurata * ''Arachniodes'' × ''mirabilis'' Sa. Kurata * ''Arachniodes'' × ''mitsuyoshiana'' Sa. Kurata * ''Arachniodes'' × ''pseudohekiana'' Kurata * ''Arachniodes'' × ''respiciens'' Sa. Kurata * ''Arachniodes'' × ''sahashii'' Fraser-Jenk. & Kandel * ''Arachniodes'' × ''sasamotoi'' Kurata * ''Arachniodes'' × ''takayamensis'' Seriz.


References


External links

{{Taxonbar, from=Q1314666 Dryopteridaceae Fern genera Taxonomy articles created by Polbot