''Arachniodes'' is a
fern genus in the family
Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the
Pteridophyte Phylogeny Group classification of 2016 (PPG I).
[ A number of species in this genus are known as "holly ferns".
The genus ''Arachniodes'' was first published by Carl Ludwig von Blume in 1828, with the single Indonesian species ''Arachniodes aspidioides''. The genus was not widely recognized until ]Mary Douglas Tindale
Mary Douglas Tindale (19 September 1920 – 31 March 2011) was an Australian botanist specialising in pteridology (ferns) and the genera '' Acacia'' and ''Glycine''.
Tindale was born in Randwick, New South Wales, the only child of George Harold ...
transferred the two species (''Byrsopteris amabilis'' and ''Byrsopteris aristata'') into it in 1961.[Tindale, Mary Douglas; ''Contributions from the New South Wales National Herbarium'' 3(1): 90. 1961.]
Species
The Pteridophyte Phylogeny Group classification of 2016 (PPG I) considers ''Arachniodes'' to have 60 species.[ , the ''Checklist of Ferns and Lycophytes of the World'' accepted 78 species:][
*'']Arachniodes ailaoshanensis
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' Ching
*''Arachniodes amabilis
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' (Blume) Tindale
*''Arachniodes amoena
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' (Ching) Ching
*'' Arachniodes aristata'' (G.Forst.) Tindale
*''Arachniodes assamica
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' (Kuhn) Ohwi
*''Arachniodes bella
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' (C.Chr.) Ching
*''Arachniodes blinii
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ' ...
'' (H.Lév.) Nakaike
*''Arachniodes carvifolia
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' (Kunze) Ching
*''Arachniodes caudata
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' Ching
*''Arachniodes cavaleriei
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ' ...
'' (Christ) Ohwi
*''Arachniodes chinensis
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ' ...
'' (Rosenst.) Ching
*'' Arachniodes coniifolia'' (T.Moore) Ching
*''Arachniodes cornucervi
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' (D.Don) Fraser-Jenk.
*''Arachniodes daklakensis
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' Li Bing Zhang, N.T.Lu & X.M.Zhou
*''Arachniodes davalliaeformis
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' (Christ) Nakaike
*'' Arachniodes denticulata'' (Sw.) Ching
*'' Arachniodes fengii'' Ching
*'' Arachniodes festina'' (Hance) Ching
*'' Arachniodes formosa'' (Fée) Ching
*'' Arachniodes formosissima'' (Goldm.) Nakaike
*'' Arachniodes gigantea'' Ching
*'' Arachniodes globisora'' (Hayata) Ching
*'' Arachniodes grossa'' (Tardieu & C.Chr.) Ching
*'' Arachniodes hainanensis'' (Ching) Ching
*'' Arachniodes haniffii'' (Holttum) Ching
*'' Arachniodes hehaii'' Li Bing Zhang, N.T.Lu & X.F.Gao
*''Arachniodes hekiana
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' Kurata
*''Arachniodes henryi
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' (Christ) Ching
*''Arachniodes hiugana
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ' ...
'' Sa. Kurata
*''Arachniodes hunanensis
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ' ...
'' Ching
*'' Arachniodes insularis'' W.H.Wagner
*''Arachniodes japonica
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' (Kurata) Nakaike
*''Arachniodes jinpingensis
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' Y.T.Hsieh
*''Arachniodes leucostegioides
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' (C.Chr.) Ching
*'' Arachniodes longicaudata'' Li Bing Zhang, N.T.Lu & Liang Zhang
*''Arachniodes longipinna
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' Ching
*''Arachniodes microlepioides
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ' ...
'' (C.Chr.) comb. ined.
*''Arachniodes miqueliana
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' (Maxim. ex Franch. & Sav.) Ohwi
*''Arachniodes miyakei
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' (H.Itô) Shimura ex Nakaike
*'' Arachniodes mutica'' (Franch. & Sav.) Ohwi
*''Arachniodes neopodophylla
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' (Ching) Nakaike
*'' Arachniodes nigrospinosa'' (Ching) Ching
*''Arachniodes nipponica
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' (Rosenst.) Ohwi
*'' Arachniodes oosorae'' H.Itô
*'' Arachniodes palmipes'' (Kunze) Fraser-Jenk.
*''Arachniodes pseudoaristata
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' (Tagawa) Ohwi
*''Arachniodes pseudoassamica
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genu ...
'' Ching
*''Arachniodes pseudorepens
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genu ...
'' Nakaike
*''Arachniodes puncticulata
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' (Alderw.) Ching
*''Arachniodes quadripinnata
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' (Hayata) Seriz.
*''Arachniodes quangnamensis
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ' ...
'' Li Bing Zhang, N.T.Lu & Liang Zhang
*''Arachniodes repens
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genu ...
'' Kurata
*''Arachniodes rhomboidea
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' (Schott) Ching
*''Arachniodes rigidissima
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' (Hook.) Ching
*''Arachniodes sarasiniorum
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' (Christ) Nakaike
*''Arachniodes simplicior
''Arachniodes simplicior'', the simpler East Indian hollyfern, East Indian holly fern, or variegated shield fern, is a species of fern in the family Dryopteridaceae.
Distribution
China (Anhui, Chongqing, Fujian, SE-Gansu, Guangxi, Guizhou, Henan ...
'' (Makino) Ohwi
*''Arachniodes simulans
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' (Ching) Ching
*''Arachniodes sinomiqueliana
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' (Ching) Ohwi
*''Arachniodes sinorhomboidea
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ' ...
'' Ching
*''Arachniodes sledgei
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ' ...
'' Fraser-Jenk.
*''Arachniodes spectabilis
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' (Ching) Ching
*''Arachniodes squamulosa
''Arachniodes squamulosa'' is a species of fern in the family Dryopteridaceae. It is endemic to Ecuador. Its natural habitats are subtropical or tropical moist lowland forests and subtropical or tropical moist montane forests. It is threatened b ...
'' R.C.Moran & B. Øllg.
*''Arachniodes standishii
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ' ...
'' (T.Moore) Ohwi
*''Arachniodes subamabilis
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' Kurata
*''Arachniodes superba
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ' ...
'' Fraser-Jenk.
*''Arachniodes tomitae
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' Sa. Kurata
*''Arachniodes tonkinensis
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' (Ching) Ching
*''Arachniodes tripinnata
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genu ...
'' (Goldm.) Sledge
*''Arachniodes tsiangiana
''Arachniodes tsiangiana'' is a species of fern in the family Dryopteridaceae. It is endemic to China. Its natural habitat is subtropical or tropical moist lowland forests. It is threatened by habitat loss
Habitat destruction (also termed ...
'' (Ching) Nakaike
*''Arachniodes webbiana
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' (A.Braun) Schelpe
*'' Arachniodes wulingshanensis'' S.F.Wu
*''Arachniodes yakusimensis
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ...
'' (H.Itô) Nakaike
*''Arachniodes ziyunshanensis
''Arachniodes'' is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".
The genus ' ...
'' Y.T.Xie
Some hybrids were also accepted:[
* ''Arachniodes'' × ''azuminoensis'' Fujiw., Y.Matsuda, Yu. Abe & Otsuka
* ''Arachniodes'' × ''chibaensis'' Yashiro
* ''Arachniodes'' × ''clivorum'' Kurata
* ''Arachniodes'' × ''ikeminensis'' Seriz.
* ''Arachniodes'' × ''kenzo-satakei'' (Kurata) Kurata
* ''Arachniodes'' × ''kurosawae'' Shimura & Kurata
* ''Arachniodes'' × ''masakii'' Kurata
* ''Arachniodes'' × ''minamitanii'' Sa. Kurata
* ''Arachniodes'' × ''mirabilis'' Sa. Kurata
* ''Arachniodes'' × ''mitsuyoshiana'' Sa. Kurata
* ''Arachniodes'' × ''pseudohekiana'' Kurata
* ''Arachniodes'' × ''respiciens'' Sa. Kurata
* ''Arachniodes'' × ''sahashii'' Fraser-Jenk. & Kandel
* ''Arachniodes'' × ''sasamotoi'' Kurata
* ''Arachniodes'' × ''takayamensis'' Seriz.
]
References
External links
{{Taxonbar, from=Q1314666
Dryopteridaceae
Fern genera
Taxonomy articles created by Polbot