Apocrita is a
suborder of
insect
Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three ...
s in the order
Hymenoptera. It includes
wasp
A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder. ...
s,
bees, and
ant
Ants are eusocial insects of the family Formicidae and, along with the related wasps and bees, belong to the order Hymenoptera. Ants evolved from vespoid wasp ancestors in the Cretaceous period. More than 13,800 of an estimated total of 22 ...
s, and consists of many families. It contains the most advanced hymenopterans and is distinguished from
Symphyta
Sawflies are the insects of the suborder Symphyta within the order Hymenoptera, alongside ants, bees, and wasps. The common name comes from the saw-like appearance of the ovipositor, which the females use to cut into the plants where they lay ...
by the narrow "waist" (
petiole) formed between the first two segments of the actual
abdomen
The abdomen (colloquially called the belly, tummy, midriff, tucky or stomach) is the part of the body between the thorax (chest) and pelvis, in humans and in other vertebrates. The abdomen is the front part of the abdominal segment of the torso. ...
; the first abdominal segment is fused to the thorax, and is called the
propodeum
The propodeum or propodium is the first abdominal segment in Apocrita Hymenoptera (wasps, bees and ants). It is fused with the thorax to form the mesosoma. It is a single large sclerite
A sclerite (Greek , ', meaning " hard") is a hardened bod ...
. Therefore, it is general practice, when discussing the body of an apocritan in a technical sense, to refer to the
mesosoma
The mesosoma is the middle part of the body, or tagma, of arthropods whose body is composed of three parts, the other two being the prosoma and the metasoma. It bears the legs, and, in the case of winged insects, the wings.
In hymenopterans of ...
and
metasoma
The metasoma is the posterior part of the body, or tagma, of arthropods whose body is composed of three parts, the other two being the prosoma and the mesosoma. In insects, it contains most of the digestive tract, respiratory system, and circul ...
(or
gaster) rather than the "thorax" and "abdomen", respectively. The evolution of a constricted waist was an important adaption for the
parasitoid
In evolutionary ecology, a parasitoid is an organism that lives in close association with its host (biology), host at the host's expense, eventually resulting in the death of the host. Parasitoidism is one of six major evolutionarily stable str ...
lifestyle of the ancestral apocritan, allowing more maneuverability of the female's
ovipositor
The ovipositor is a tube-like organ used by some animals, especially insects, for the laying of eggs. In insects, an ovipositor consists of a maximum of three pairs of appendages. The details and morphology of the ovipositor vary, but typical ...
. The
ovipositor
The ovipositor is a tube-like organ used by some animals, especially insects, for the laying of eggs. In insects, an ovipositor consists of a maximum of three pairs of appendages. The details and morphology of the ovipositor vary, but typical ...
either extends freely or is retracted, and may be developed into a stinger for both defense and paralyzing prey. Larvae are legless and blind, and either feed inside a host (plant or animal) or in a nest cell provisioned by their mothers.
Apocrita has historically been split into two groups,
Parasitica
Parasitica (the parasitican wasps) is an obsolete, paraphyletic infraorder of Apocrita containing the parasitoid wasps. It includes all Apocrita except for the Aculeata. Parasitica has more members as a group than both the Symphyta and the Acule ...
and
Aculeata
Aculeata is a subclade of Hymenoptera containing ants, bees, and stinging wasps. The name is a reference to the defining feature of the group, which is the modification of the ovipositor into a stinger. However, many members of the group cannot ...
. Aculeata is a
clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
whose name is in standard use. "Parasitica" is not a clade, as it is
paraphyletic
In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...
: the clade would contain the Aculeata. "Parasitica" is therefore a rankless grouping in many present classifications, if it appears at all.
Parasitica comprises the majority of hymenopteran insects, its members living as
parasitoids
In evolutionary ecology, a parasitoid is an organism that lives in close association with its host at the host's expense, eventually resulting in the death of the host. Parasitoidism is one of six major evolutionary strategies within parasi ...
.
Most species are small, with the ovipositor adapted for piercing. In some hosts, the parasitoids induce metamorphosis
Metamorphosis is a biological process by which an animal physically develops including birth or hatching, involving a conspicuous and relatively abrupt change in the animal's body structure through cell growth and differentiation. Some inse ...
prematurely, and in others it is prolonged. There are even species that are hyperparasite
A hyperparasite, also known as a metaparasite, is a parasite whose host, often an insect, is also a parasite, often specifically a parasitoid. Hyperparasites are found mainly among the wasp-waisted Apocrita within the Hymenoptera, and in two oth ...
s, or parasitoids on other parasitoids. The Parasitica lay their egg
An egg is an organic vessel grown by an animal to carry a possibly fertilized egg cell (a zygote) and to incubate from it an embryo within the egg until the embryo has become an animal fetus that can survive on its own, at which point the a ...
s inside or on another insect (egg, larva or pupa) and their larvae grow and develop within or on that host. The host is nearly always killed. Many parasitic hymenopterans are used as biological control
Biological control or biocontrol is a method of controlling pests, such as insects, mites, weeds, and plant diseases, using other organisms. It relies on predation, parasitism, herbivory, or other natural mechanisms, but typically also invo ...
agents to control pests, such as caterpillar
Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths).
As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder Sym ...
s, true bugs and hoppers, flies
Flies are insects of the order Diptera, the name being derived from the Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwings having evolved into advanced ...
, and weevil
Weevils are beetles belonging to the Taxonomic rank, superfamily Curculionoidea, known for their elongated snouts. They are usually small, less than in length, and Herbivore, herbivorous. Approximately 97,000 species of weevils are known. They b ...
s.
Aculeata is a monophyletic
In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ...
group that includes those species in which the female's ovipositor
The ovipositor is a tube-like organ used by some animals, especially insects, for the laying of eggs. In insects, an ovipositor consists of a maximum of three pairs of appendages. The details and morphology of the ovipositor vary, but typical ...
is modified into a stinger
A stinger (or sting) is a sharp organ found in various animals (typically insects and other arthropods) capable of injecting venom, usually by piercing the epidermis of another animal.
An insect sting is complicated by its introduction of v ...
to inject venom
Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
. Groups within Aculeata include the familiar ants, bees, and various types of parasitic and predatory wasps; it also includes all of the social
Social organisms, including human(s), live collectively in interacting populations. This interaction is considered social whether they are aware of it or not, and whether the exchange is voluntary or not.
Etymology
The word "social" derives from ...
hymenopterans.
Among the nonparasitic and nonsocial Aculeata, larvae are fed with captured prey (typically alive and paralyzed) or may be fed pollen and nectar. The social
Social organisms, including human(s), live collectively in interacting populations. This interaction is considered social whether they are aware of it or not, and whether the exchange is voluntary or not.
Etymology
The word "social" derives from ...
Aculeata feed their young prey (paper wasp
Paper wasps are Vespidae, vespid wasps that gather fibers from dead wood and plant stems, which they mix with saliva, and use to construct nests made of gray or brown papery material. Some types of paper wasps are also sometimes called umbrell ...
s and hornet
Hornets (insects in the genus ''Vespa'') are the largest of the eusocial wasps, and are similar in appearance to their close relatives yellowjackets. Some species can reach up to in length. They are distinguished from other vespine wasps by th ...
s), or pollen
Pollen is a powdery substance produced by seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametophyt ...
and nectar
Nectar is a sugar-rich liquid produced by plants in glands called nectaries or nectarines, either within the flowers with which it attracts pollinating animals, or by extrafloral nectaries, which provide a nutrient source to animal mutualists ...
(bees), or perhaps seeds, fungi
A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from ...
, or nonviable eggs (ants).
Extant families and superfamilies
The Apocrita contains a large number of families. Some traditional taxa such as the Parasitica
Parasitica (the parasitican wasps) is an obsolete, paraphyletic infraorder of Apocrita containing the parasitoid wasps. It includes all Apocrita except for the Aculeata. Parasitica has more members as a group than both the Symphyta and the Acule ...
(containing many families of parasitoid wasp
Parasitoid wasps are a large group of hymenopteran superfamilies, with all but the wood wasps (Orussoidea) being in the wasp-waisted Apocrita. As parasitoids, they lay their eggs on or in the bodies of other arthropods, sooner or later causin ...
s) have been found on molecular analysis to be paraphyletic
In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...
. Parasitoid
In evolutionary ecology, a parasitoid is an organism that lives in close association with its host (biology), host at the host's expense, eventually resulting in the death of the host. Parasitoidism is one of six major evolutionarily stable str ...
ism evolved once, and it is found today across most Apocritan families, though it has been secondarily lost several times. The phylogenetic tree
A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spec ...
gives a condensed overview of the phylogeny, illustrated with major groups. The sawflies are paraphyletic as the Apocrita evolved inside that group. The tree is not fully resolved.
*Suborder Apocrita
**Infraorder Aculeata
Aculeata is a subclade of Hymenoptera containing ants, bees, and stinging wasps. The name is a reference to the defining feature of the group, which is the modification of the ovipositor into a stinger. However, many members of the group cannot ...
***Superfamily Apoidea
The superfamily (zoology), superfamily Apoidea is a major group within the Hymenoptera, which includes two traditionally recognized lineages, the "sphecidae, sphecoid" wasps, and the bees. Molecular phylogeny demonstrates that the bees arose from ...
( bees and sphecoid wasps)
****Family Ampulicidae ( cockroach wasps)
****Family Andrenidae
The Andrenidae (commonly known as mining bees) are a large, nearly cosmopolitan family of solitary, ground-nesting bees. Most of the family's diversity is located in temperate or arid areas (warm temperate xeric). It includes some enormous gener ...
(mining bees)
****Family Apidae
Apidae is the largest family within the superfamily Apoidea, containing at least 5700 species of bees. The family includes some of the most commonly seen bees, including bumblebees and honey bees, but also includes stingless bees (also used for ...
(carpenter bee
Carpenter bees are species in the genus ''Xylocopa'' of the subfamily Xylocopinae. The genus includes some 500 bees in 31 subgenera. The common name "carpenter bee" derives from their nesting behavior; nearly all species burrow into hard plant m ...
s, digger bee
The Anthophorini are a large tribe in the subfamily Apinae of the family Apidae. Species in this tribe are often referred to as digger bees, although this common name is sometimes also applied to members of the tribe Centridini. It contains over ...
s, cuckoo bee
The term cuckoo bee is used for a variety of different bee lineages which have evolved the kleptoparasitism, kleptoparasitic behaviour of laying their eggs in the nests of other bees, reminiscent of the behavior of cuckoo birds. The name is perhap ...
s, bumble bee
A bumblebee (or bumble bee, bumble-bee, or humble-bee) is any of over 250 species in the genus ''Bombus'', part of Apidae, one of the bee families. This genus is the only Extant taxon, extant group in the tribe Bombini, though a few extinct r ...
s, orchid bee
The tribe Euglossini, in the subfamily Apinae, commonly known as orchid bees or euglossine bees, are the only group of corbiculate bees whose non-parasitic members do not all possess eusocial behavior.
Description
Most of the tribe's species ...
s, stingless bee
Stingless bees, sometimes called stingless honey bees or simply meliponines, are a large group of bees (about 550 described species), comprising the tribe Meliponini (or subtribe Meliponina according to other authors). They belong in the family A ...
s, and honeybee
A honey bee (also spelled honeybee) is a eusocial flying insect within the genus ''Apis'' of the bee clade, all native to Afro-Eurasia. After bees spread naturally throughout Africa and Eurasia, humans became responsible for the current cosmo ...
s)
****Family Colletidae
The Colletidae are a family of bees, and are often referred to collectively as plasterer bees or polyester bees, due to the method of smoothing the walls of their nest cells with secretions applied with their mouthparts; these secretions dry into ...
( yellow-faced bees and plasterer bee
A plasterer is a tradesman or tradesperson who works with plaster, such as forming a layer of plaster on an interior wall or plaster decorative moldings on ceilings or walls. The process of creating plasterwork, called plastering, has been ...
s)
****Family Crabronidae
The Crabronidae are a large paraphyletic group (nominally a family) of wasps, including nearly all of the species formerly comprising the now-defunct superfamily Sphecoidea. It collectively includes well over 200 genera, containing well over 9 ...
(sand wasps, bee wolves, etc.)
****Family Halictidae
Halictidae is the second-largest family of bees (clade Anthophila) with nearly 4,500 species. Halictid species are an extremely diverse group that can vary greatly in appearance. These bees occur all over the world and are found on every contine ...
("sweat bees")
****Family Heterogynaidae
****Family Megachilidae
Megachilidae is a cosmopolitan family of mostly solitary bees. Both that their pollen-carrying structure (called a ''scopa'') is restricted to the ventral surface of the abdomen (rather than mostly or exclusively on the hind legs as in other b ...
( leaf-cutting bees)
****Family Melittidae
Melittidae is a small bee family, with over 200 described species in three subfamilies. The family has a limited distribution, with all described species restricted to Africa and the northern temperate zone.
Fossil melittids have been found oc ...
****Family Stenotritidae
The Stenotritidae is the smallest of all formally recognised bee families, with only 21 species in two genera, all of them restricted to Australia. Historically, they were generally considered to belong in the family Colletidae, but the stenotrit ...
****Family Sphecidae ( digger wasps)
***Superfamily Chrysidoidea
The superfamily Chrysidoidea is a very large cosmopolitan group (some 6,000 described species, and many more undescribed) , all of which are parasitoids or cleptoparasites of other insects. There are three large, common families (Bethylidae, Chr ...
****Family Bethylidae
The Bethylidae are a family of aculeate wasps in the superfamily Chrysidoidea. As a family, their biology ranges between parasitoid wasps and hunting wasps.
Overview
Like most of the Chrysidoidea, the Bethylidae are stinging Hymenoptera and m ...
****Family Chrysididae
Commonly known as cuckoo wasps or emerald wasps, the hymenopteran family Chrysididae is a very large cosmopolitan group (over 3000 described species) of parasitoid or kleptoparasitic wasps, often highly sculptured, with brilliant metallic colors ...
(cuckoo wasp
Commonly known as cuckoo wasps or emerald wasps, the hymenopteran family Chrysididae is a very large cosmopolitan group (over 3000 described species) of parasitoid or kleptoparasitic wasps, often highly sculptured, with brilliant metallic colors ...
s)
****Family Dryinidae
Dryinidae is a cosmopolitan family of solitary wasps. Its name comes from the Greek ''drys'' for oak: Latreille named the type genus ''Dryinus'' because the first species was collected in an oak plant in Spain. The larvae are parasitoids of the ...
****Family Embolemidae
Embolemidae is a family of small solitary parasitoid wasps with around 70 species in 2 genera distributed around the world.van Achterberg, Cornelis & Kats, R.. (2000). Revision of the Palaearctic Embolemidae (Hymenoptera). Zoöl. Med. 74 (2000), ...
****Family Plumariidae
****Family Sclerogibbidae
The Sclerogibbidae are a small family of aculeate wasps in the superfamily Chrysidoidea.
Overview
Sclerogibbidae are ectoparasitoids of Embioptera. The female wasp oviposits an egg on the abdomen of a host. Once the larva emerges, it attaches i ...
****Family Scolebythidae
The Scolebythidae are a small family of aculeate wasps in the superfamily Chrysidoidea. These chrysidoid wasps are found in Africa, Australia, the Neotropics, north China, Thailand and Fiji. They are parasites on larvae of Cerambycidae and Ptini ...
***Superfamily Formicoidea
Ants are eusocial insects of the family Formicidae and, along with the related wasps and bees, belong to the order Hymenoptera. Ants evolved from vespoid wasp ancestors in the Cretaceous period. More than 13,800 of an estimated total of 22,00 ...
****Family Formicidae (ant
Ants are eusocial insects of the family Formicidae and, along with the related wasps and bees, belong to the order Hymenoptera. Ants evolved from vespoid wasp ancestors in the Cretaceous period. More than 13,800 of an estimated total of 22 ...
s)
***Superfamily Pompiloidea
Pompiloidea is a superfamily that includes spider wasps and velvet ants, among others. in the order Hymenoptera. There are 4 families in Pompiloidea.
Families
These four families belong to the superfamily Pompiloidea:
* Mutillidae (velvet ant ...
****Family Mutillidae
The Mutillidae are a family of more than 7,000 species of wasps whose wingless females resemble large, hairy ants. Their common name velvet ant refers to their dense pile of hair, which most often is bright scarlet or orange, but may also be ...
(velvet ant
The Mutillidae are a family of more than 7,000 species of wasps whose wingless females resemble large, hairy ants. Their common name velvet ant refers to their dense pile of hair, which most often is bright scarlet or orange, but may also be bl ...
s)
****Family Myrmosidae
The Myrmosidae are a small family of wasps very similar to the Mutillidae. As in mutillids, females are flightless, and are kleptoparasites in the nests of fossorial bees and wasps.
Taxonomy
Recent classifications of Vespoidea ''sensu lato'' (be ...
****Family Pompilidae
Wasps in the family Pompilidae are commonly called spider wasps, spider-hunting wasps, or pompilid wasps. The family is cosmopolitan, with some 5,000 species in six subfamilies. Nearly all species are solitary (with the exception of some group-ne ...
(spider wasp
Wasps in the family Pompilidae are commonly called spider wasps, spider-hunting wasps, or pompilid wasps. The family is cosmopolitan, with some 5,000 species in six subfamilies. Nearly all species are solitary (with the exception of some group-ne ...
s)
****Family Sapygidae
The Sapygidae are a family of solitary kleptoparasitic aculeate wasps. They are generally black wasps, similar in appearance to some Tiphiidae or Thynnidae, with white or yellow markings developed to various degrees.
The female oviposits her egg ...
***Superfamily Scolioidea
The Scoliidae, the scoliid wasps, are a family of about 560 species found worldwide. They tend to be black, often marked with yellow or orange, and their wing tips are distinctively corrugated. Males are more slender and elongated than females, wi ...
****Family Scoliidae
The Scoliidae, the scoliid wasps, are a family of about 560 species found worldwide. They tend to be black, often marked with yellow or orange, and their wing tips are distinctively corrugated. Males are more slender and elongated than females, wi ...
***Superfamily Tiphioidea
Tiphioidea is a suggested superfamily of stinging wasps in the order Hymenoptera. There are three families in Tiphioidea, Bradynobaenidae, Tiphiidae, and Sierolomorphidae.
Recent research in molecular phylogenetics has resulted in the reorgani ...
****Family Bradynobaenidae
The Bradynobaenidae are a family of wasps similar to the Mutillidae, differing most visibly in the presence, in females, of a suture separating the pronotum from the mesonotum. These species are often found in arid regions. Recent classifications ...
****Family Sierolomorphidae
The Sierolomorphidae are a family of 13 extant species of wasps, in the genera ''Sierolomorpha'' and ''Proscleroderma'', mostly found in the Northern Hemisphere. They are rare and very little is known of their biology. A fossil species ''Loreisom ...
****Family Tiphiidae
The Tiphiidae (also known as the tiphiid wasps) are a family of large, solitary wasps whose larvae are parasitoids of various beetle larvae, especially those in the superfamily Scarabaeoidea. Until recently, this family contained several addition ...
***Superfamily Thynoidea
****Family Chyphotidae
The Chyphotidae are a family of wasps similar to the Mutillidae, differing most visibly in the presence, in females, of a suture separating the pronotum from the mesonotum. These species are found primarily in arid regions in the southwestern Uni ...
****Family Thynnidae
The Thynnidae (also known as thynnid wasps or flower wasps) are a family of large, solitary wasps whose larvae are almost universally parasitoids of various beetle larvae, especially those in the superfamily Scarabaeoidea. Until recently, the co ...
***Superfamily Vespoidea
Vespoidea is a superfamily of wasps in the order Hymenoptera, although older taxonomic schemes may vary in this categorization, particularly in whether to recognize the superfamilies Scolioidea (for scoliid wasps) or Formicoidea (for ants). V ...
****Family Rhopalosomatidae
Rhopalosomatidae is a family of Hymenoptera containing about 68 extant species in four genera that are found worldwide. Three fossil genera are known.
The adults resemble ants and may be confused with them. They are yellowish with red or brown m ...
****Family Vespidae
The Vespidae are a large (nearly 5000 species), diverse, cosmopolitan family of wasps, including nearly all the known eusocial wasps (such as ''Polistes fuscatus'', ''Vespa orientalis'', and ''Vespula germanica'') and many solitary wasps. Each ...
(paper wasp
Paper wasps are Vespidae, vespid wasps that gather fibers from dead wood and plant stems, which they mix with saliva, and use to construct nests made of gray or brown papery material. Some types of paper wasps are also sometimes called umbrell ...
s, potter wasp
Potter wasps (or mason wasps), the Eumeninae, are a cosmopolitan wasp group presently treated as a subfamily of Vespidae, but sometimes recognized in the past as a separate family, Eumenidae.
Recognition
Most eumenine species are black or brown, ...
s, hornet
Hornets (insects in the genus ''Vespa'') are the largest of the eusocial wasps, and are similar in appearance to their close relatives yellowjackets. Some species can reach up to in length. They are distinguished from other vespine wasps by th ...
s, pollen wasps, yellowjacket
Yellowjacket or yellowjacket is the common name in North America for predatory social wasps of the genus, genera ''Vespula'' and ''Dolichovespula''. Members of these genera are known simply as "wasps" in other English-speaking countries. Most of ...
s)
**Infraorder Parasitica
Parasitica (the parasitican wasps) is an obsolete, paraphyletic infraorder of Apocrita containing the parasitoid wasps. It includes all Apocrita except for the Aculeata. Parasitica has more members as a group than both the Symphyta and the Acule ...
***Superfamily Ceraphronoidea
****Family Ceraphronidae
The Ceraphronidae are a small hymenopteran family with 14 genera and some 360 known species, though a great many species are still undescribed. It is a poorly known group as a whole, though most are believed to be parasitoids (especially of f ...
****Family Megaspilidae
The Megaspilidae are a small hymenopteran family with 13 genera in two subfamilies, and some 450 known species, with a great many species still undescribed. It is a poorly known group as a whole, though most are believed to be parasitoids (espe ...
***Superfamily Chalcidoidea
Chalcid wasps (, , for their metallic colour) are insects within the superfamily Chalcidoidea, part of the order Hymenoptera. The superfamily contains some 22,500 known species, and an estimated total diversity of more than 500,000 species, me ...
****Family Agaonidae
The family Agaonidae is a group of pollinating and nonpollinating fig wasps. They spend their larval stage inside the fruits of figs. The pollinating wasps (Agaoninae, Kradibiinae, and Tetrapusiinae) are the mutualistic partners of the fig trees ...
(fig wasp
Fig wasps are wasps of the superfamily Chalcidoidea which spend their larval stage inside figs. Most are pollinators but others simply feed off the plant. The non-pollinators belong to several groups within the superfamily Chalcidoidea, while the ...
s)
****Family Aphelinidae
****Family Chalcididae
The Chalcididae are a moderate-sized family within the Chalcidoidea, composed mostly of parasitoids and a few hyperparasitoids. The family is apparently polyphyletic, though the different subfamilies may each be monophyletic, and some may be elev ...
( chalcid wasps)
****Family Encyrtidae
Encyrtidae is a large family of parasitic wasps, with some 3710 described species in about 455 genera. The larvae of the majority are primary parasitoids on Hemiptera, though other hosts are attacked, and details of the life history can be variab ...
****Family Eucharitidae
The Eucharitidae are a family of parasitic wasps.Ayre, G.L. Pseudometagea schwarzii (Ashm.) (Eucharitidae: Hymenoptera), a parasite of Lasius neoniger Emery (Formicidae: Hymenoptera). Canadian Journal of Zoology 40 (1962) : 157-164. Eucharitid ...
****Family Eulophidae
The Eulophidae are a large family of hymenopteran insects, with over 4,300 described species in some 300 genera. The family includes the genus '' Elasmus'', which used to be treated as a separate family, "Elasmidae", and is now treated as a s ...
****Family Eupelmidae
Eupelmidae is a family of parasitic wasps in the superfamily Chalcidoidea. The group is apparently polyphyletic, though the different subfamilies may each be monophyletic, and may be elevated to family status in the near future. As presently defi ...
****Family Eurytomidae ( seed chalcids)
****Family Leucospidae
****Family Mymaridae
The Mymaridae, commonly known as fairyflies or fairy wasps, are a family of chalcidoid wasps found in temperate and tropical regions throughout the world. The family contains around 100 genera with 1400 species.
Fairyflies are very tiny insects ...
(fairyflies
The Mymaridae, commonly known as fairyflies or fairy wasps, are a family of chalcidoid wasps found in temperate and tropical regions throughout the world. The family contains around 100 genera with 1400 species.
Fairyflies are very tiny insects ...
) – the smallest of all insects
****Family Ormyridae
The Ormyridae are a small family of parasitic wasps in the superfamily Chalcidoidea. They are either parasitoids or hyperparasitoids on gall-forming insects, primarily cynipid wasps and tephritid flies. The 120 or so species in three genera (m ...
****Family Perilampidae
The Perilampidae are a small family within the Chalcidoidea, composed mostly of hyperparasitoids. The family is closely related to the Eucharitidae, and the eucharitids appear to have evolved from within the Perilampidae, thus rendering the fa ...
****Family Pteromalidae
The Pteromalidae are a very large family of mostly parasitoid wasps, with some 3,450 described species in about 640 genera (the number was greater, but many species and genera have been reduced by synonymy in recent years). The subfamily-level d ...
****Family Rotoitidae
****Family Signiphoridae
****Family Tanaostigmatidae
****Family Tetracampidae
****Family Torymidae
Torymidae is a family of wasps in the superfamily Chalcidoidea. Most species in this family are small with attractive metallic coloration, and females generally have long ovipositors. Many are parasitoids on gall-forming insects, and some are phy ...
****Family Trichogrammatidae
The Trichogrammatidae are a family of tiny wasps in the Chalcidoidea that include some of the smallest of all insects, with most species having adults less than 1 mm in length, with species of '' Megaphragma'' having an adult body length ...
***Superfamily Cynipoidea
The Cynipoidea are a moderate-sized hymenopteran superfamily that presently includes five modern families and three extinct families, though others have been recognized in the past. The most familiar members of the group are phytophagous, especia ...
****Family Austrocynipidae
****Family Cynipidae
Gall wasps, also incorrectly called gallflies, are hymenopterans of the family Cynipidae in the wasp superfamily Cynipoidea. Their common name comes from the galls they induce on plants for larval development. About 1,300 species of this genera ...
(gall wasp
Gall wasps, also incorrectly called gallflies, are hymenopterans of the family Cynipidae in the wasp superfamily Cynipoidea. Their common name comes from the galls they induce on plants for larval development. About 1,300 species of this generall ...
s)
****Family Figitidae
Figitidae is a family of parasitoid wasps. The full diversity of this wasp family is not yet known, but about 1400 species have been described to over 130 genera. For example, the largest subfamily, Eucoilinae (previously considered as a separ ...
****Family Ibaliidae
The Ibaliidae are a small family of hymenopteran superfamily Cynipoidea. Ibaliidae differ from most of the cynipoids by the larvae being parasitoids on other wasp larvae in the group Siricidae. The Ibaliidae comprise three extant genera of f ...
****Family Liopteridae
Liopteridae is a family of wood-boring parasitoid wasp
Parasitoid wasps are a large group of hymenopteran superfamilies, with all but the wood wasps (Orussoidea) being in the wasp-waisted Apocrita. As parasitoids, they lay their eggs on o ...
***Superfamily Diaprioidea
****Family Austroniidae
****Family Diapriidae
The Diapriidae are a family of parasitoid wasps. These tiny insects have an average length of 2–4 mm and never exceed 8 mm. They typically attack larvae and pupae of a wide range of insects, especially flies. The about 2,300 described ...
****Family Maamingidae
''Maaminga '' is a genus of parasitic wasps containing two species and constituting the family Maamingidae. So far only known from New Zealand, the two named species are '' Maaminga marrisi'' and '' M. rangi''. The genus (and family) name derive ...
****Family Monomachidae
***Superfamily Evanioidea
****Family Aulacidae
The Aulacidae are a small, cosmopolitan family of wasps, with two extant genera containing some 200 known species. They are primarily endoparasitoids of wood wasps (Xiphydriidae) and xylophagous beetles (Cerambycidae and Buprestidae). They are c ...
****Family Evaniidae
Evaniidae is a family of parasitoid wasps also known as ensign wasps, nightshade wasps, hatchet wasps, or cockroach egg parasitoid wasps. They number around 20 extant genera containing over 400 described species, and are found all over the world e ...
( ensign wasps)
****Family Gasteruptiidae
The Gasteruptiidae are one of the more distinctive families among the apocritan wasps, with surprisingly little variation in appearance for a group that contains around 500 species in two subfamilies ( Gasteruptiinae and Hyptiogastrinae) and wi ...
***Superfamily Ichneumonoidea
The superfamily Ichneumonoidea contains one extinct and three extant families, including the two largest families within Hymenoptera: Ichneumonidae and Braconidae. The group is thought to contain as many as 100,000 species, many of which have not ...
****Family Braconidae
The Braconidae are a family of parasitoid wasps. After the closely related Ichneumonidae, braconids make up the second-largest family in the order Hymenoptera, with about 17,000 recognized species and many thousands more undescribed. One analysis ...
****Family Ichneumonidae
The Ichneumonidae, also known as the ichneumon wasps, Darwin wasps, or ichneumonids, are a family (biology), family of parasitoid wasps of the insect order Hymenoptera. They are one of the most diverse groups within the Hymenoptera with roughly 2 ...
(ichneumon wasp
The Ichneumonidae, also known as the ichneumon wasps, Darwin wasps, or ichneumonids, are a family of parasitoid wasps of the insect order Hymenoptera. They are one of the most diverse groups within the Hymenoptera with roughly 25,000 species cu ...
s)
***Superfamily Megalyroidea
Megalyroidea is a small hymenopteran superfamily of wasps that includes a single family, Megalyridae, with eight extant genera (plus around a dozen extinct ones) and 49 described species. Modern megalyrids are found primarily in the southern hem ...
****Family Megalyridae
***Superfamily Mymarommatoidea
The Mymarommatoidea are a very small superfamily of microscopic fairyfly-like parasitic wasps. It contains only a single living family, Mymarommatidae, and three other extinct families known from Cretaceous aged amber. Less than half of all descr ...
– sometimes called Serphitoidea
****Family Mymarommatidae
The Mymarommatidae, sometimes referred to as false fairy wasps. are a very small family of microscopic parasitic wasps. Only about half of the known species are living taxa (the others are fossils), but they are found worldwide.Gibson, G.A.P.; Re ...
***Superfamily Platygastroidea
The Hymenopteran superfamily of parasitoid wasps, Platygastroidea, has often been treated as a lineage within the superfamily Proctotrupoidea, but most classifications since 1977 have recognized it as an independent group within the Proctotrupo ...
****Family Platygastridae
The hymenopteran family Platygastridae (sometimes incorrectly spelled Platygasteridae) is a large group (over 4000 species) of exclusively parasitoid wasps, mostly very small (1–2 mm), black, and shining, with geniculate (elbowed) antennae ...
****Family Scelionidae
The hymenopteran family Scelionidae is a very large cosmopolitan group (over 3000 described species in some 176 genera) of exclusively parasitoid wasps, mostly small (0.5–10 mm), often black, often highly sculptured, with (typically) elbowe ...
***Superfamily Proctotrupoidea
Proctotrupoidea is a hymenopteran superfamily containing seven extant families, though others have been recognized in the past, most of these having been removed to a recently erected superfamily Diaprioidea. Of the remaining families, only Pr ...
****Family Heloridae
Heloridae is a family of wasps in the order Hymenoptera known primarily from fossils, and only one extant genus, '' Helorus'', with 12 species found worldwide. Members of ''Helorus'' are parasitic on green lacewings.
Extant genera
*'' Helorus'' ( ...
****Family Pelecinidae
Pelecinidae is a family of parasitic wasps in the superfamily Proctotrupoidea. It contains only one living genus, '' Pelecinus'', with three species known from the Americas. The earliest fossil species are known from the Jurassic, and the group ...
****Family Peradeniidae
****Family Proctorenyxidae
****Family Proctotrupidae
****Family Roproniidae
****Family Vanhorniidae
Vanhorniidae is a family parasitic wasps belonging to the Proctotrupoidea. There are only two extant genera, '' Heloriserphus'' and '' Vanhornia,'' which are native to the Northern Hemisphere. They are parasitoids of beetle larvae belonging to the ...
***Superfamily Stephanoidea
****Family Stephanidae
***Superfamily Trigonaloidea
****Family †Maimetshidae
Maimetshidae is an extinct family of wasps, known from the Cretaceous period.M. S. Engel. 2016A new genus and species of maimetshid wasps in Lebanese Early Cretaceous amber (Hymenoptera: Maimetshidae).''Novitates Paleoentomologicae'' 18:1-14 While ...
****Family Trigonalidae
Trigonalidae (the spelling Trigonalyidae is incorrect under Article 29.5 of the ICZN,[ICZN Code](_blank)
29.5. Maintena ...
References
External links
Suborder Apocrita – Ants, Bees and Wasps – BugGuide.Net
— images and other information
Tree of Life
Balades Entomologiques
— "entomological walks" with images
{{Taxonbar, from=Q22864
Insect suborders
Taxa named by Carl Eduard Adolph Gerstaecker