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The rock cale (''Aplodactylus lophodon''), also known as the cockatoo fish, cocky, joey, rock cocky or sea carp, is a species of marine ray finned fish, one of the marblefishes belonging to the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Aplodactylidae. It is found in the southwestern Pacific Ocean.


Taxonomy

The rock cale was first formally described in 1859 by the
German German(s) may refer to: * Germany (of or related to) **Germania (historical use) * Germans, citizens of Germany, people of German ancestry, or native speakers of the German language ** For citizens of Germany, see also German nationality law **Ger ...
born British herpetologist and
ichthyologist Ichthyology is the branch of zoology devoted to the study of fish, including bony fish ( Osteichthyes), cartilaginous fish (Chondrichthyes), and jawless fish (Agnatha). According to FishBase, 33,400 species of fish had been described as of Octobe ...
Albert Günther Albert Karl Ludwig Gotthilf Günther FRS, also Albert Charles Lewis Gotthilf Günther (3 October 1830 – 1 February 1914), was a German-born British zoologist, ichthyologist, and herpetologist. Günther is ranked the second-most productive re ...
, although he misspelt the genus name as ''Haplodactylus'', with the type locality given as
New South Wales ) , nickname = , image_map = New South Wales in Australia.svg , map_caption = Location of New South Wales in AustraliaCoordinates: , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , es ...
. For some time some workers considered that this species was in its own monospecific
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''Crinodus'' but in 2000 Barry C. Russell of the
Museum and Art Gallery of the Northern Territory The Museum and Art Gallery of the Northern Territory (MAGNT) is the main museum in the Northern Territory. The museum is located in the inner Darwin suburb of Fannie Bay. The MAGNT is governed by the Board of the Museum and Art Gallery of the ...
reviewed the family Apodactylidae and returned this species to the genus ''Aplodactylus''. The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
''lophodon'' is a compound of ''lophos'' which means "tuft" and ''odon'' meaning "tooth", a reference to the terminal lobe and two lateral lobes on each side of its incisor like teeth.


Description

The rock cale has an elongate body which has a depth equivalent to roughly a third to a quarter of its
standard length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish m ...
. It has a short snout and a rounded head which has a small mouth which is slightly downturned and has fleshy lips, the upper lip projecting. The jaws have small, The jaws have small, tricuspid teeth with a few sharply pointed ones too and they are set on 3-4 rows in the jaws. The outermost row has the largest teeth and there are no
vomerine teeth The vomer (; lat, vomer, lit=ploughshare) is one of the unpaired facial bones of the skull. It is located in the midsagittal line, and articulates with the sphenoid, the ethmoid, the left and right palatine bones, and the left and right maxill ...
. There are two pairs of nostrils, the front pair have fleshy flap with small tentacles on the front and rear margins. There is a wide, flattened spine in the operculum which does not reach the margin. The
dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through conv ...
has a long base but it is nearly split in two by a deep and wide incision between the spiny and soft rayed parts. The dorsal fin contains 17 spines and 18-20 soft rays while the
anal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
has 3 spines and 6-7 soft rays. It has large pectoral fins which may be longer than the head and the lowest 5-6 rays are unbranched and fleshy. The body is covered in small, cycloid scales which are embedded in the skin, these extend on to the cheeks and operculum and create a sheath along base of the spiny portion of the dorsal fin. The largest
total length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish ...
recorded is . The overall colour of the rock cale varies from grey to bluish-black, broken by whitish mottles on the flanks and whitish spots on the fins. There is a dark saddle blotch underneath the middle of spiny part of the dorsal fin, and a line of five uniformly spaced white blotches on the flanks below the
lateral line The lateral line, also called the lateral line organ (LLO), is a system of sensory organs found in fish, used to detect movement, vibration, and pressure gradients in the surrounding water. The sensory ability is achieved via modified epithelial ...
, starting above the pectoral fin and ending under the rear end of the dorsal fin. The juveniles have an obvious black mark on upper rear edge of operculum.


Distribution and habitat

The rock cale is endemic to the southeastern Australian waters of the southwestern Pacific Ocean. Here it occurs from
Mudjimba Mudjimba is a coastal suburb in the Sunshine Coast Region, Queensland, Australia, and forms part of the Maroochydore urban centre. In the , Mudjimba had a population of 2,540 people. Geography The suburb is bounded by Mudjimba Beach on the ...
on the
Sunshine Coast Sunshine Coast may refer to: * Sunshine Coast, Queensland, Australia **Sunshine Coast Region, a local government area of Queensland named after the region **Sunshine Coast Stadium * Sunshine Coast (British Columbia), geographic subregion of the Br ...
of
Queensland ) , nickname = Sunshine State , image_map = Queensland in Australia.svg , map_caption = Location of Queensland in Australia , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , established_ ...
south to the
Gippsland Lakes The Gippsland Lakes are a network of coastal lakes, marshes and lagoons in East Gippsland, Victoria, Australia covering an overall area of about between the rural towns of Lakes Entrance, Bairnsdale and Sale. The largest of the lakes are La ...
in Victoria, and Babel Island off
Tasmania ) , nickname = , image_map = Tasmania in Australia.svg , map_caption = Location of Tasmania in AustraliaCoordinates: , subdivision_type = Country , subdi ...
. It lives on shallow, exposed rocky shores, schooling in high-energy zone, just below the surface down to around where there is good cover of kelp and other seaweeds.


Biology

The rock cale is herbivorous feeding on weeds and algae, such as ''
Corallina pilulifera ''Corallina'' is a genus of red seaweeds with hard, abrasive calcareous skeletons in the family Corallinaceae. They are stiff, branched plants with articulations. Species # '' Corallina aberrans'' (Yendo) K.R.Hind & G.W.Saunders # '' Coralli ...
'', as well as lesser amounts of ''
Ulva lactuca ''Ulva lactuca'', also known by the common name sea lettuce, is an edible green alga in the family Ulvaceae. It is the type species of the genus ''Ulva''. A synonym is ''U. fenestrata'', referring to its "windowed" or "holed" appearance. Descrip ...
'', '' Jania'' and filamentous algae including species in the genus '' Sphacelaria''. They will feed on both free floating and attached algae. Typically this species is observed sitting on the "turf" made up of coralline algae but when there is strong wave surges it can wedge itself into crevices in the rock. It is one of the most numerous reef fishes in the surge zone off New South Wales.


References

{{Taxonbar, from=Q6422428 Aplodactylidae Fish described in 1859 Taxa named by Albert Günther