''Anisancylus dutrae'' is a

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

of small,

freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include ...

, air-breathing

limpet Limpets are a group of aquatic snails that exhibit a conical shell shape (patelliform) and a strong, muscular foot. Limpets are members of the class Gastropoda, but are polyphyletic, meaning the various groups called "limpets" descended indep ...

, an aquatic

pulmonate Pulmonata or pulmonates, is an informal group (previously an order, and before that a subclass) of snails and slugs characterized by the ability to breathe air, by virtue of having a pallial lung instead of a gill, or gills. The group includ ...

gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. T ...

mollusc Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000 extant species of molluscs are recognized. The number of fossil species is esti ...

in the family Planorbidae, the ram's horn snails and their allies.

Etymology

The species name honours Dr. Ana Virginea C. Dutra of the

Federal University of Pernambuco Federal University of Pernambuco ( pt, Universidade Federal de Pernambuco, UFPE) is a public university in Recife, Brazil, established in 1946. UFPE has 70 undergraduate courses and 175 postgraduate courses. , UFPE had 35,000 students and 2,000 ...

Distribution

''A. dutrae'' is

endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...

Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...

, where it occurs in the northeastern states of

Pernambuco Pernambuco () is a state of Brazil, located in the Northeast region of the country. With an estimated population of 9.6 million people as of 2020, making it seventh-most populous state of Brazil and with around 98,148 km², being the 19 ...

and

Bahia Bahia ( , , ; meaning "bay") is one of the 26 Federative units of Brazil, states of Brazil, located in the Northeast Region, Brazil, Northeast Region of the country. It is the fourth-largest Brazilian state by population (after São Paulo (sta ...

. The species' type locality is the Tapacurá Ecological Station in

São Lourenço da Mata São Lourenço da Mata is a city located in the greater Recife metropolitan area in the state of Pernambuco, with a population of 114,079 inhabitants (2020 estimate by IBGE). The city was one of the hosts of the 2014 FIFA World Cup together with ...

Description

''A. dutrae'' has a high shell with an oval

aperture In optics, an aperture is a hole or an opening through which light travels. More specifically, the aperture and focal length of an optical system determine the cone angle of a bundle of rays that come to a focus in the image plane. An opt ...

, reaching in length. The apex is prominent, not pointed, dorsally flattened, bent to the right and back. The

protoconch A protoconch (meaning first or earliest or original shell) is an embryonic or larval shell which occurs in some classes of molluscs, e.g., the initial chamber of an ammonite or the larval shell of a gastropod. In older texts it is also called ...

shows an apical depression and a small flat area, followed by irregularly arranged punctuations. The radial lines are imperceptible to optical microscopy. The

teleoconch The gastropod shell is part of the body of a gastropod or snail, a kind of mollusc. The shell is an exoskeleton, which protects from predators, mechanical damage, and dehydration, but also serves for muscle attachment and calcium storage. Some ...

shows prominent radial lines across the surface of the shell and concentric, wavy fine lines. The

mantle A mantle is a piece of clothing, a type of cloak. Several other meanings are derived from that. Mantle may refer to: *Mantle (clothing), a cloak-like garment worn mainly by women as fashionable outerwear **Mantle (vesture), an Eastern Orthodox ve ...

is pigmented, generally dark brown or black, except in the areas of muscle insertion (foot and tentacles). The right anterior muscle impression is elongated, extending to the midline of the body, with a median constriction. The left impression is elliptical, transversely arranged. The posterior impression is elliptical. The

radula The radula (, ; plural radulae or radulas) is an anatomical structure used by molluscs for feeding, sometimes compared to a tongue. It is a minutely toothed, chitinous ribbon, which is typically used for scraping or cutting food before the food ...

shows marked characteristics. There is a quadricuspid rachidian tooth, the two main cusps being asymmetrical and aculeate. Beside each principal cusp there is one small accessory cusp. There are three principal cusps in the lateral teeth. The mesocone is the most developed, long and aculeate, while the endocone is relatively short. There is a big gap between the mesocone and the ectocone. The

ovotestis An ovotestis is a gonad with both testicular and ovarian aspects. In humans, ovotestes are an infrequent anatomical variation associated with gonadal dysgenesis. The only mammals where ovotestes are not symptomatic of an intersex variation are mole ...

is wide, having about 35 unbranched, closely pressed follicles. The prostate has 4-5 follicles.

Habitat

''A. dutrae'' is found attached to the upper surface of stones in

lotic River ecosystems are flowing waters that drain the landscape, and include the Biotic component, biotic (living) interactions amongst plants, animals and micro-organisms, as well as abiotic (nonliving) physical and chemical interactions of its man ...

habitats (such as small, fast-flowing streams).

References

{{Taxonbar, from=Q17080689 Planorbidae Molluscs of Brazil Gastropods described in 1994