''Animantarx'' ( ; meaning 'living citadel') is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

of nodosaurid ankylosaurian

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

from the

Early Early may refer to: History * The beginning or oldest part of a defined historical period, as opposed to middle or late periods, e.g.: ** Early Christianity ** Early modern Europe Places in the United States * Early, Iowa * Early, Texas * Early ...

and

Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...

of western

North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...

. Like other nodosaurs, it would have been a slow-moving quadrupedal

herbivore A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpart ...

covered in heavy armor scutes, but without a

tail club In zoology, a club is a bony mass at the end of the tail of some dinosaurs and of some mammals, most notably the ankylosaurids and the glyptodonts, as well as meiolaniid turtles. It is thought that this was a form of defensive armour or weapon ...

. The

skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, the ...

measures approximately 25 cm (10 inches) in length, suggesting the animal as a whole was no more than 3 meters (10 feet) long.

Discovery and species

The generic name is composed of the

Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...

words ''animatus'' ("living" or "animated") and ''arx'' ("fortress" or "citadel"), referring to its armored nature. In particular, the name is a reference to a comment made by

paleontologist Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fossi ...

R. S. Lull Richard Swann Lull (November 6, 1867 – April 22, 1957) was an American paleontologist and Sterling Professor at Yale University who is largely remembered now for championing a non-Darwinian view of evolution, whereby mutation(s) could unlo ...

about ankylosaurs, that as "an animated citadel, these animals must have been practically unassailable..." The

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

is the only one known so far, and is called ''A. ramaljonesi'' after its discoverer, Ramal Jones. His wife, Carol Jones, also discovered the contemporaneous dinosaur '' Eolambia'' nearby. Only one specimen of ''Animantarx'' has so far been recovered. The remains include the lower jaw and back half of the skull, along with neck and back

vertebra The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic ...

e, and various limb elements. ''Animantarx'' is characterized by a unique combination of features, including a highly domed skull back, small horns on the postorbital and quadratojugal bones of the skull, and a

mandible In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower tooth, teeth in place. The mandible sits beneath the maxilla. It is the only movabl ...

which is only armoured on half of its length. These

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

remains were discovered in the Mussentuchit Member of the Cedar Mountain Formation in the eastern portion of the U.S. state of

Utah Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to it ...

. This section of the

formation Formation may refer to: Linguistics * Back-formation, the process of creating a new lexeme by removing or affixes * Word formation, the creation of a new word by adding affixes Mathematics and science * Cave formation or speleothem, a secondar ...

is believed to represent the late Albian through early

Cenomanian The Cenomanian is, in the ICS' geological timescale, the oldest or earliest age of the Late Cretaceous Epoch or the lowest stage of the Upper Cretaceous Series. An age is a unit of geochronology; it is a unit of time; the stage is a unit in the s ...

stages of the

Period, or about 106 to 97 million years ago. At least 80 other

vertebrate Vertebrates () comprise all animal taxa within the subphylum Vertebrata () ( chordates with backbones), including all mammals, birds, reptiles, amphibians, and fish. Vertebrates represent the overwhelming majority of the phylum Chordata, ...

species are known from the Mussentuchit, including

fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...

frog A frog is any member of a diverse and largely Carnivore, carnivorous group of short-bodied, tailless amphibians composing the order (biology), order Anura (ανοὐρά, literally ''without tail'' in Ancient Greek). The oldest fossil "proto-f ...

lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia alt ...

snake Snakes are elongated, Limbless vertebrate, limbless, carnivore, carnivorous reptiles of the suborder Serpentes . Like all other Squamata, squamates, snakes are ectothermic, amniote vertebrates covered in overlapping Scale (zoology), scales. Ma ...

crocodilia Crocodilia (or Crocodylia, both ) is an order of mostly large, predatory, semiaquatic reptiles, known as crocodilians. They first appeared 95 million years ago in the Late Cretaceous period ( Cenomanian stage) and are the closest living ...

ns,

bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...

s, and

mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...

s, although not all are complete enough to name. Many dinosaur groups are represented by fossils from this

member Member may refer to: * Military jury, referred to as "Members" in military jargon * Element (mathematics), an object that belongs to a mathematical set * In object-oriented programming, a member of a class ** Field (computer science), entries in ...

, including

carnivorous A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose food and energy requirements derive from animal tissues (mainly muscle, fat and other sof ...

theropods as well as several different herbivorous types, including the iguanodont ''Eolambia''. The presence of aquatic animals like fish and frogs, as well as the

mudstone Mudstone, a type of mudrock, is a fine-grained sedimentary rock whose original constituents were clays or muds. Mudstone is distinguished from '' shale'' by its lack of fissility (parallel layering).Blatt, H., and R.J. Tracy, 1996, ''Petrology. ...

in which their fossils are found, suggests that this was a

floodplain A floodplain or flood plain or bottomlands is an area of land adjacent to a river which stretches from the banks of its channel to the base of the enclosing valley walls, and which experiences flooding during periods of high discharge.Goudi ...

environment.Kirkland, J.I., Britt, B., Burge, D.L., Carpenter, K., Cifelli, R., DeCourten, F., Eaton, J., Hasiotis, S., and Lawton, T. 1997. Lower to Middle Cretaceous dinosaur faunas of the Central Colorado Plateau: a key to understanding 35 million years of tectonics, sedimentology, evolution, and biogeography. ''Brigham Young University Geology Studies'' 42:69-103. Earlier layers within the Cedar Mountain Formation contain different nodosaur species. The oldest layer, known as the Yellow Cat Member, contains '' Gastonia'', while the intermediate Poison Strip and Ruby Ranch Members contain remains which may belong to ''

Sauropelta ''Sauropelta'' ( ; meaning 'lizard shield') is a genus of nodosaurid dinosaur that existed in the Early Cretaceous Period of North America. One species (''S. edwardsorum'') has been named although others may have existed. Anatomically, ''Saurop ...

''. The Mussentuchit, which is the youngest member of the Cedar Mountain, contains only ''Animantarx''. While there is still a lot of exploration left to be done, this division of nodosaur species corresponds with that of other dinosaur groups and provides support for the

hypothesis A hypothesis (plural hypotheses) is a proposed explanation for a phenomenon. For a hypothesis to be a scientific hypothesis, the scientific method requires that one can test it. Scientists generally base scientific hypotheses on previous obse ...

of three separate

fauna Fauna is all of the animal life present in a particular region or time. The corresponding term for plants is ''flora'', and for fungi, it is '' funga''. Flora, fauna, funga and other forms of life are collectively referred to as '' biota''. Zoo ...

s in the Cedar Mountain Formation. The Mussentuchit fauna includes many taxa which may be of

Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...

n origin and suggests a dispersal event may have occurred from Asia into North America around this time. Fossils in this region are often slightly

radioactive Radioactive decay (also known as nuclear decay, radioactivity, radioactive disintegration, or nuclear disintegration) is the process by which an unstable atomic nucleus loses energy by radiation. A material containing unstable nuclei is consid ...

, and remains of ''Animantarx'' were actually discovered following a radiological survey of the area performed by Ramal Jones, which located a higher level of radioactivity at a certain location. Subsequent excavation at this site turned up the fossil skeleton of ''Animantarx''; no bones had been exposed on the surface.

Classification

''Animantarx'' is universally thought of as a nodosaurid ankylosaur, although its precise relationships within that family are uncertain. The most recent cladistic analysis of ankylosaur phylogeny does not include ''Animantarx'', although the authors recognize the genus as Nodosauridae '' incertae sedis'' because of its rounded

supraorbital Supraorbital refers to the region immediately above the eye sockets, where in humans the eyebrows are located. It denotes several anatomical features, such as: *Supraorbital artery *Supraorbital foramen *Supraorbital gland *Supraorbital nerve *Sup ...

protrusions and a "knoblike"

acromion In human anatomy, the acromion (from Greek: ''akros'', "highest", ''ōmos'', "shoulder", plural: acromia) is a bony process on the scapula (shoulder blade). Together with the coracoid process it extends laterally over the shoulder joint. The acro ...

on the

scapula The scapula (plural scapulae or scapulas), also known as the shoulder blade, is the bone that connects the humerus (upper arm bone) with the clavicle (collar bone). Like their connected bones, the scapulae are paired, with each scapula on eithe ...

. Two separate studies have found ''Animantarx'' to be the sister taxon of '' Edmontonia'' within Nodosauridae.Hill, R.V., Witmer, L.M., Norell, M.A. 2003. A New specimen of ''Pinacosaurus grangeri'' (Dinosauria: Ornithischia) from the Late Cretaceous of Mongolia: ontogeny and phylogeny of ankylosaurs. ''American Museum Novitates'' 3395: 1-29.

References

{{Taxonbar, from=Q132966 Nodosaurids Cretaceous dinosaurs of North America Albian life Cenomanian life Fossil taxa described in 1999 Taxa named by James I. Kirkland Taxa named by Kenneth Carpenter Ornithischian genera

Discovery and species

Classification

See also

References