Animal Feces
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Feces ( or faeces), known colloquially and in slang as poo and poop, are the solid or semi-solid remains of food that was not digested in the
small intestine The small intestine or small bowel is an organ in the gastrointestinal tract where most of the absorption of nutrients from food takes place. It lies between the stomach and large intestine, and receives bile and pancreatic juice through the p ...
, and has been broken down by bacteria in the large intestine. Feces contain a relatively small amount of
metabolic waste Metabolic wastes or excrements are substances left over from metabolic processes (such as cellular respiration) which cannot be used by the organism (they are surplus or toxic), and must therefore be excreted. This includes nitrogen compounds, ...
products such as bacterially altered
bilirubin Bilirubin (BR) (Latin for "red bile") is a red-orange compound that occurs in the normal catabolic pathway that breaks down heme in vertebrates. This catabolism is a necessary process in the body's clearance of waste products that arise from the ...
, and dead epithelial cells from the lining of the gut. Feces are discharged through the
anus The anus (Latin, 'ring' or 'circle') is an opening at the opposite end of an animal's digestive tract from the mouth. Its function is to control the expulsion of feces, the residual semi-solid waste that remains after food digestion, which, d ...
or
cloaca In animal anatomy, a cloaca ( ), plural cloacae ( or ), is the posterior orifice that serves as the only opening for the digestive, reproductive, and urinary tracts (if present) of many vertebrate animals. All amphibians, reptiles and birds, a ...
during
defecation Defecation (or defaecation) follows digestion, and is a necessary process by which organisms eliminate a solid, semisolid, or liquid waste material known as feces from the digestive tract via the anus. The act has a variety of names ranging f ...
. Feces can be used as fertilizer or
soil conditioner A soil conditioner is a product which is added to soil to improve the soil’s physical qualities, usually its fertility (ability to provide nutrition for plants) and sometimes its mechanics. In general usage, the term "soil conditioner" is often ...
in agriculture. They can also be burned as
fuel A fuel is any material that can be made to react with other substances so that it releases energy as thermal energy or to be used for work. The concept was originally applied solely to those materials capable of releasing chemical energy but ...
or dried and used for construction. Some medicinal uses have been found. In the case of
human feces Human feces (or faeces in British English) is the solid or semisolid remains of food that could not be digested or absorbed in the small intestine of humans, but has been further broken down by bacteria in the large intestine. It also contains b ...
, fecal transplants or fecal bacteriotherapy are in use. Urine and feces together are called excreta. Skatole is the principal compound responsible for the unpleasant smell of feces.


Characteristics

The distinctive
odor An odor (American English) or odour (English in the Commonwealth of Nations, Commonwealth English; American and British English spelling differences#-our, -or, see spelling differences) is caused by one or more volatilized chemical compounds ...
of feces is due to
skatole Skatole or 3-methylindole is an organic compound belonging to the indole family. It occurs naturally in the feces of mammals and birds and is the primary contributor to fecal odor. In low concentrations, it has a flowery smell and is found in seve ...
, and thiols (
sulfur Sulfur (or sulphur in British English) is a chemical element with the symbol S and atomic number 16. It is abundant, multivalent and nonmetallic. Under normal conditions, sulfur atoms form cyclic octatomic molecules with a chemical formula ...
-containing compounds), as well as amines and carboxylic acids. Skatole is produced from tryptophan via indoleacetic acid.
Decarboxylation Decarboxylation is a chemical reaction that removes a carboxyl group and releases carbon dioxide (CO2). Usually, decarboxylation refers to a reaction of carboxylic acids, removing a carbon atom from a carbon chain. The reverse process, which is t ...
gives skatole. The perceived bad odor of feces has been hypothesized to be a deterrent for humans, as consuming or touching it may result in sickness or infection.


Physiology

Feces are discharged through the
anus The anus (Latin, 'ring' or 'circle') is an opening at the opposite end of an animal's digestive tract from the mouth. Its function is to control the expulsion of feces, the residual semi-solid waste that remains after food digestion, which, d ...
or
cloaca In animal anatomy, a cloaca ( ), plural cloacae ( or ), is the posterior orifice that serves as the only opening for the digestive, reproductive, and urinary tracts (if present) of many vertebrate animals. All amphibians, reptiles and birds, a ...
during
defecation Defecation (or defaecation) follows digestion, and is a necessary process by which organisms eliminate a solid, semisolid, or liquid waste material known as feces from the digestive tract via the anus. The act has a variety of names ranging f ...
. This process requires pressures that may reach (13.3 kPa) in humans and (60 kPa) in penguins. The forces required to expel the feces is generated through muscular contractions and a build-up of gases inside the gut, prompting the sphincter to relieve the pressure and to release the feces.


Ecology

After an animal has digested eaten material, the remains of that material are discharged from its body as waste. Although it is lower in energy than the food from which it is derived, feces may retain a large amount of energy, often 50% of that of the original food. This means that of all food eaten, a significant amount of energy remains for the decomposers of ecosystems. Many organisms feed on feces, from bacteria to fungi to insects such as dung beetles, who can sense odors from long distances. Some may specialize in feces, while others may eat other foods. Feces serve not only as a basic food, but also as a supplement to the usual diet of some animals. This process is known as
coprophagia Coprophagia () or coprophagy () is the consumption of feces. The word is derived from the grc, κόπρος , "feces" and , "to eat". Coprophagy refers to many kinds of feces-eating, including eating feces of other species (heterospecifics), o ...
, and occurs in various animal species such as young elephants eating the feces of their mothers to gain essential gut flora, or by other animals such as dogs, rabbits, and monkeys. Feces and urine, which reflect ultraviolet light, are important to raptors such as kestrels, who can see the near ultraviolet and thus find their prey by their middens and territorial markers. Seeds also may be found in feces. Animals who eat fruit are known as frugivores. An advantage for a plant in having fruit is that animals will eat the fruit and unknowingly disperse the seed in doing so. This mode of seed dispersal is highly successful, as seeds dispersed around the base of a plant are unlikely to succeed and often are subject to heavy seed predation, predation. Provided the seed can withstand the pathway through the digestive system, it is not only likely to be far away from the parent plant, but is even provided with its own fertilizer. Organisms that subsist on dead organic matter or ''detritus'' are known as detritivores, and play an important role in ecosystems by recycling organic matter back into a simpler form that plants and other autotrophs may absorb once again. This cycling of matter is known as the biogeochemical cycle. To maintain nutrients in soil it is therefore important that feces returns to the area from which they came, which is not always the case in human society where food may be transported from rural areas to urban populations and then feces disposed of into a river or sea.


Human feces

Depending on the individual and the circumstances, human beings may defecate several times a day, every day, or once every two or three days. Extensive hardening of the feces that interrupts this routine for several days or more is called constipation. The appearance of human fecal matter varies according to diet and health. Normally it is semisolid, with a mucus coating. A combination of bile and
bilirubin Bilirubin (BR) (Latin for "red bile") is a red-orange compound that occurs in the normal catabolic pathway that breaks down heme in vertebrates. This catabolism is a necessary process in the body's clearance of waste products that arise from the ...
, which comes from dead red blood cells, gives feces the typical brown color. After the meconium, the first stool expelled, a newborn's feces contains only bile, which gives it a yellow-green color. Breast feeding babies expel soft, pale yellowish, and not quite malodorous matter; but once the baby begins to eat, and the body starts expelling bilirubin from dead red blood cells, its matter acquires the familiar brown color. At different times in their life, human beings will expel feces of different colors and textures. A stool that passes rapidly through the intestines will look greenish; lack of bilirubin will make the stool look like clay.


Uses of animal feces


Fertilizer

The feces of animals, e.g. guano and manure often are used as fertilizer.


Energy

Dry animal dung fuel, Dry animal dung, such as that of camel, bison and cattle, is burned as fuel many countries. Animals such as the giant panda and zebra possess gut bacteria capable of producing biofuel. The bacterium in question, Brocadia anammoxidans, can be used to synthesize the rocket fuel hydrazine.


Coprolites and paleofeces

A coprolite is fossilized feces and is classified as a trace fossil. In paleontology they give evidence about the diet of an animal. They were first described by William Buckland in 1829. Prior to this, they were known as "fossil conifer cone#Pinaceae cones, fir cones" and "bezoar stones". They serve a valuable purpose in paleontology because they provide direct evidence of the predation and diet of extinct organisms. Coprolites may range in size from a few millimetres to more than 60 centimetres. Palaeofeces are ancient human feces, feces, often found as part of archaeology, archaeological excavations or surveys. Intact paleofeces of ancient people may be found in caves in arid climates and in other locations with suitable preservation conditions. These are studied to determine the diet (nutrition), diet and health of the people who produced them through the analysis of seeds, small bones, and parasite eggs found inside. Feces may contain information about the person excreting the material as well as information about the material. They also may be analyzed chemically for more in-depth information on the individual who excreted them, using lipid analysis and ancient DNA analysis. The success rate of usable DNA extraction is relatively high in paleofeces, making it more reliable than skeletal DNA retrieval. The reason this analysis is possible at all is due to the digestive system not being entirely efficient, in the sense that not everything that passes through the digestive system is destroyed. Not all of the surviving material is recognizable, but some of it is. Generally, this material is the best indicator archaeologists can use to determine ancient diets, as no other part of the archaeological record is so direct an indicator. A process that preserves feces in a way that they may be analyzed later is the Maillard reaction. This reaction creates a casing of sugar that preserves the feces from the elements. To extract and analyze the information contained within, researchers generally have to freeze the feces and grind it up into powder for analysis.


Other uses

Animal dung occasionally is used as a cement to make adobe mudbrick huts, or even in throwing sports, especially with cow and camel dung. ''Kopi luwak'' (pronounced ), or "civet coffee", is coffee made from coffee berries that have been eaten by and passed through the digestive tract of the Asian palm civet (''Paradoxurus hermaphroditus''). Giant pandas provide fertilizer for the world's most expensive green tea. In Malaysia, tea is made from the droppings of stick insects fed on guava leaves. In northern Thailand, elephants are used to digest coffee beans in order to make Black Ivory coffee, which is among the world's most expensive coffees. Paper is also made from elephant dung in the country. Dog feces was used in the Tanning (leather), tanning process of leather during the Victorian era. Collected dog feces, known as "pure", "puer", or "pewer", was mixed with water to form a substance known as "bate", because proteolytic enzymes in the dog feces helped to relax the fibrous structure of the hide before the final stages of tanning. Dog feces collectors were known as ''pure finders''. Elephants, hippopotamus, hippos, koalas and pandas are born with sterile intestines, and require bacteria obtained from eating the feces of their mothers to digest vegetation. In India, cow dung and cow urine are major ingredients of the traditional Hindu drink ''Panchagavya''. Politician Shankarbhai Vegad stated that they can cure cancer. In the Middle East, cow dung is consumed for a variety of reasons, such as curing dysentery, a belief of healing properties or as a food staple.


Terminology

Feces is the scientific terminology, while the term stool is also commonly used in medical contexts. Outside of scientific contexts, these terms are less common, with the most common layman's term being "poop" or "poo". The term shit is also in common use, although is widely considered vulgar or offensive. There are many other terms, see below.


Etymology

The word ''faeces'' is the plural of the Latin word '':wikt:faex, faex'' meaning "dregs". In most English-language usage, there is no singular form, making the word a plurale tantum; out of various major dictionaries, only one enters English plurals#Plural in form but singular in construction, variation from plural agreement.


Synonyms

"Feces" is used more in biology and medicine than in other fields (reflecting science's tradition of classical Latin and New Latin) *In hunting and tracking, terms such as ''dung'', ''scat'', ''spoor'', and ''droppings'' normally are used to refer to non-human animal feces *In husbandry and farming, ''manure'' is common. *''Stool'' is a common term in reference to
human feces Human feces (or faeces in British English) is the solid or semisolid remains of food that could not be digested or absorbed in the small intestine of humans, but has been further broken down by bacteria in the large intestine. It also contains b ...
. For example, in medicine, to diagnose the presence or absence of a medical condition, a stool sample sometimes is requested for testing purposes. *The term ''bowel movement(s)'' (with each movement a defecation event) is also common in health care. There are many synonyms in informal register (sociolinguistics), registers for feces, just like there are for urine. Many are euphemismistic, colloquialism, colloquial, or both; some are profanity, profane (such as ''shit''), whereas most belong chiefly to baby talk, child-directed speech (such as ''poo'' or the palindromic, palindromic word ''poop'') or to off-color humor, crude humor (such as ''crap, dump, load'' and ''turd''.).


Feces of animals

The feces of animals often has special names (some of them are slang), for example: *Non-human animals **As bulk material – dung **Individually – droppings *Cattle **Bulk material – cow dung **Individual droppings – cow pats, meadow muffins, etc. *Deer (and formerly other quarry animals) – fewmets * Wild Carnivora, carnivores – scat *Otter – spraint *Birds (individual) – droppings (also include urine as white crystals of uric acid) *Seabirds or bats (large accumulations) – guano *Herbivorous insects, such as caterpillars and leaf beetles – frass *Earthworms, lugworms etc. – worm castings (feces extruded at ground surface) *Feces when used as fertilizer (usually mixed with animal bedding and urine) – manure *Horses – horse manure, roadapple (before motor vehicles became common, horse droppings were a big part of the rubbish communities needed to clean off roads)


Society and culture


Feelings of disgust

In all human cultures, feces elicits varying degrees of disgust in adults. Children under two years typically have no disgust response to it, suggesting it is culturally derived. Disgust toward feces appears to be strongest in cultures where flush toilets make olfactory contact with human feces minimal. Disgust is experienced primarily in relation to the sense of taste (either perceived or imagined) and, secondarily to anything that causes a similar feeling by sense of smell, touch, or vision.


Social media

There is a Pile of Poo emoji represented in Unicode as , called ' or ''unchi-kun'' in Japan.


Jokes

Poop is the center of Toilet humour, toilet humor, and is commonly in interest of young children and teenagers.


Gallery

File:BearApplePoop.JPG, Bear scat File:Mixedplasticbearpoo.jpg, Bear scat showing consumption of bin bags File:Kasuar fg1.jpg, The cassowary disperses plant seeds via its feces File:Earthworm faeces.jpg, Earthworm feces aids in provision of minerals and plant nutrients in an accessible form File:Avföring - Fågelspillning - Ystad-2015.jpg, Feces from different seabirds.


See also

* Coprophagia * Coprophilia * Coprophilous fungi * Cow dung * Excretory system * Fecal plug * Jenkem * Manure * Metabolic waste * Night soil * Sanitation * Scatology * Whale feces * Zooplankton fecal pellets


References


External links


MedFriendly's Article on Feces
{{Authority control Feces, Animal physiology