''Anemone hepatica'' (

syn. The Botanical and Zoological Codes of nomenclature treat the concept of synonymy differently. * In botanical nomenclature, a synonym is a scientific name that applies to a taxon that (now) goes by a different scientific name. For example, Linnae ...

''Hepatica nobilis''), the common hepatica, liverwort, kidneywort, or pennywort, is a

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...

in the buttercup

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Ranunculaceae Ranunculaceae (buttercup or crowfoot family; Latin "little frog", from "frog") is a family of over 2,000 known species of flowering plants in 43 genera, distributed worldwide. The largest genera are ''Ranunculus'' (600 species), ''Delphinium' ...

, native to woodland in

temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ...

regions of the Northern Hemisphere. This

herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition of t ...

perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...

grows from a

rhizome In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow hori ...

Description

Hepatica nobilis flowers - blue and pink - Keila

''Anemone hepatica'' grows high. Leaves and flowers emerge directly from the

, not from a stem above ground. The leaves have three lobes and are fleshy and hairless, wide and long . The upper side is dark green with whitish stripes and the lower side is violet or reddish brown. Leaves emerge during or after flowering and remain green through winter. The flowers are blue, purple, pink, or white and appear in winter or spring. They have five to ten oval showy

sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined b ...

s and three green

bract In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves. They may be smaller, larger, or of ...

Taxonomy

The taxonomy of the genus ''Anemone'' and its species is not fully resolved, but

phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...

studies of many species of ''Anemone'' and related genera indicate that species of the genus ''

Hepatica ''Hepatica'' (hepatica, liverleaf, or liverwort) is a genus of herbaceous perennials in the buttercup family, native to central and northern Europe, Asia and eastern North America. Some botanists include ''Hepatica'' within a wider interpre ...

'' should be included under ''Anemone'' because of similarities both in molecular attributes and other shared morphologies. The circumscription of the taxon is also debated, some authors listing the North American var. ''acuta'' and var. ''obtusa'', while other list them as the separate species ''A. acutiloba'' and ''A. americana'', respectively.

Varieties

Varieties of ''Anemone hepatica'' that are sometimes recognized include: * ''Anemone hepatica'' var. ''japonica'', a synonym of ''Hepatica nobilis'' var. ''japonica'' , is native to the

Russian Far East The Russian Far East (russian: Дальний Восток России, r=Dal'niy Vostok Rossii, p=ˈdalʲnʲɪj vɐˈstok rɐˈsʲiɪ) is a region in Northeast Asia. It is the easternmost part of Russia and the Asian continent; and is admini ...

China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...

Korea Korea ( ko, 한국, or , ) is a peninsular region in East Asia. Since 1945, it has been divided at or near the 38th parallel, with North Korea (Democratic People's Republic of Korea) comprising its northern half and South Korea (Republic o ...

, and

Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...

* ''Anemone hepatica'' var. ''acuta'', a synonym of '' Hepatica acutiloba'' , is native to eastern

North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...

* ''Anemone hepatica'' var. ''obtusa'', a synonym of ''

Hepatica americana ''Hepatica americana'', the round-lobed hepatica, is a herbaceous flowering plant in the buttercup family Ranunculaceae. It is native to the eastern United States and Canada. It is sometimes considered part of the genus ''Anemone'', as ''Anemo ...

'' , is native to eastern North America Asian liverleaf.jpg, ''Anemone hepatica'' var. ''japonica''

Distribution and habitat

It is found in woods, thickets and meadows, especially in the mountains of continental Europe, North America and Japan.

Ecology

Hepatica flowers produce pollen but no nectar. In North America, the flowers first attract ''

Lasioglossum The sweat bee genus ''Lasioglossum'' is the largest of all bee genera, containing over 1700 species in numerous subgenera worldwide.Gibbs, J., et al. (2012)Phylogeny of halictine bees supports a shared origin of eusociality for ''Halictus'' an ...

'' sweat bees and

small carpenter bees The cosmopolitan bee genus ''Ceratina'', often referred to as small carpenter bees, is the sole lineage of the tribe Ceratinini, and is not closely related to the more familiar carpenter bees. The genus presently contains over 300 species in 23 ...

looking in vain for nectar. Then when the

stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...

s begin to release pollen, the bees return to collect and feed on pollen.

Mining bee ''Andrena'' is a genus of bees in the family Andrenidae. With over 1,500 species, it is one of the largest genera of animals. It is a strongly monophyletic group that is difficult to split into more manageable divisions; currently, ''Andrena'' i ...

s sometimes visit the flowers, but prefer flowers that produce both nectar and pollen.

Toxicity

Like other Ranunculaceae, fresh liverwort contains

protoanemonin Protoanemonin (sometimes called anemonol or ranunculol) is a toxin found in all plants of the buttercup family (Ranunculaceae). When the plant is wounded or macerated, the unstable glucoside found in the plant, ranunculin, is enzymatically broken d ...

and is therefore slightly toxic. By drying the herb, protoanemonin is

dimerize A dimer () ('' di-'', "two" + ''-mer'', "parts") is an oligomer consisting of two monomers joined by bonds that can be either strong or weak, covalent or intermolecular. Dimers also have significant implications in polymer chemistry, inorganic ch ...

d to the non-toxic

anemonin Anemonin is a compound found in plants of the buttercup family (Ranunculaceae). It is the dimerization product of the toxin protoanemonin and easily reacts with water to a dicarboxylic acid. The name of the substance comes from the plant gen ...

Uses

Medieval

herbalist Herbal medicine (also herbalism) is the study of pharmacognosy and the use of medicinal plants, which are a basis of traditional medicine. With worldwide research into pharmacology, some herbal medicines have been translated into modern remed ...

s believed it could be used to treat liver diseases, and is still used in alternative medicine today. Other modern applications by herbalists include treatments for

pimple A pimple is a kind of comedo that results from excess sebum and dead skin cells getting trapped in the pores of the skin. In its aggravated state, it may evolve into a pustule or papules. Pimples can be treated by acne medications, antibiotics, ...

bronchitis Bronchitis is inflammation of the bronchi (large and medium-sized airways) in the lungs that causes coughing. Bronchitis usually begins as an infection in the nose, ears, throat, or sinuses. The infection then makes its way down to the bronchi. ...

and

gout Gout ( ) is a form of inflammatory arthritis characterized by recurrent attacks of a red, tender, hot and swollen joint, caused by deposition of monosodium urate monohydrate crystals. Pain typically comes on rapidly, reaching maximal intensit ...

. Under the name ''Hepatica nobilis'', which is now regarded as a synonym, this plant has gained the

Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr (Nort ...

Award of Garden Merit The Award of Garden Merit (AGM) is a long-established annual award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. History The Award of Garden Merit ...

Culture

It is the official flower of the

Sweden Democrats The Sweden Democrats ( sv , Sverigedemokraterna ; SD ) is a nationalist and right-wing populist political party in Sweden. As of 2022, it is the largest member of Sweden's right-wing governing bloc to which it provides confidence and supply, a ...

political party in Swedish politics.

Gallery

File:Ranuncolaceae - Anemone hepatica-2.JPG File:Anemone hepatica in the Horsan nature reserve.jpg File:Ranuncolaceae - Anemone hepatica.JPG File:Sinilill, makro.jpg File:Hepatica nobilis Kelchblätter W.jpg File:Hepatica nobilis kz1.jpg

References

External links

*
Flora of Pennsylvania
{{Taxonbar, from1=Q15613598, from2=Q161050

hepatica ''Hepatica'' (hepatica, liverleaf, or liverwort) is a genus of herbaceous perennials in the buttercup family, native to central and northern Europe, Asia and eastern North America. Some botanists include ''Hepatica'' within a wider interpre ...

Medicinal plants of Europe Medicinal plants of North America Flora of Europe Flora of Asia Plants described in 1753 Taxa named by Carl Linnaeus Flora of Eastern Canada Flora of the Northeastern United States Flora of the North-Central United States Flora of the Southeastern United States Flora without expected TNC conservation status