The clouded salamander (''Aneides ferreus'') is a species of

salamander Salamanders are a group of amphibians typically characterized by their lizard-like appearance, with slender bodies, blunt snouts, short limbs projecting at right angles to the body, and the presence of a tail in both larvae and adults. All t ...

in the family

Plethodontidae Plethodontidae, or lungless salamanders, are a family of salamanders. Most species are native to the Western Hemisphere, from British Columbia to Brazil, although a few species are found in Sardinia, Europe south of the Alps, and South Korea. In ...

. It is

endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...

to the

Pacific Northwest The Pacific Northwest (sometimes Cascadia, or simply abbreviated as PNW) is a geographic region in western North America bounded by its coastal waters of the Pacific Ocean to the west and, loosely, by the Rocky Mountains to the east. Tho ...

. Its natural

habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...

is temperate

forest A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...

s and it is probable that many nest in trees. It is threatened by

habitat loss Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...

Description

The clouded salamander can grow to about total length. It is long and slim with relatively long legs, square ends to its toes and a prehensile tail. It has two naso-labial grooves joining its nostrils to its mouth and sixteen costal grooves down its flank. The upper side is pale grey variously blotched with gold, olive green or dull red. The juvenile has a brass-colored streak down the center of its back.

Distribution and habitat

The clouded salamander is found in the Cascadian bioregion of North America at heights up to . Its range extends from the Columbia River southwards through the

Cascade Mountains The Cascade Range or Cascades is a major mountain range of western North America, extending from southern British Columbia through Washington and Oregon to Northern California. It includes both non-volcanic mountains, such as the North Cascades, a ...

, along the Oregon coast to the northern tip of California in

Del Norte County ), in California , seat_type = County seat , seat = Crescent City , parts_type = Largest city , parts = Crescent City , unit_pref = US , area_total_sq_mi = 1230 , area_land_sq_mi = 1006 , area_water_sq_mi = 223 , elevation_max_footnote ...

. It lives in forested areas favoring moist areas with few trees rather than dry dense stands. It hides under logs and rocks, under loose bark, in rotting logs and in crevices in rock and scree. It particularly favors rotting

Douglas-fir The Douglas fir (''Pseudotsuga menziesii'') is an evergreen conifer species in the pine family, Pinaceae. It is native to western North America and is also known as Douglas-fir, Douglas spruce, Oregon pine, and Columbian pine. There are three va ...

logs. It is often common in newly cleared areas with tree stumps and debris.

Biology

The clouded salamander feeds on small invertebrates, mainly

woodlice A woodlouse (plural woodlice) is an isopod crustacean from the polyphyleticThe current consensus is that Oniscidea is actually triphyletic suborder Oniscidea within the order Isopoda. They get their name from often being found in old wood ...

ant Ants are eusocial insects of the family Formicidae and, along with the related wasps and bees, belong to the order Hymenoptera. Ants evolved from vespoid wasp ancestors in the Cretaceous period. More than 13,800 of an estimated total of ...

s and

beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...

s, but also include

flies Flies are insects of the order Diptera, the name being derived from the Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwings having evolved into advanced m ...

termite Termites are small insects that live in colonies and have distinct castes (eusocial) and feed on wood or other dead plant matter. Termites comprise the infraorder Isoptera, or alternatively the epifamily Termitoidae, within the order Blatto ...

mite Mites are small arachnids (eight-legged arthropods). Mites span two large orders of arachnids, the Acariformes and the Parasitiformes, which were historically grouped together in the subclass Acari, but genetic analysis does not show clear e ...

s, centipedes,

millipede Millipedes are a group of arthropods that are characterised by having two pairs of jointed legs on most body segments; they are known scientifically as the class Diplopoda, the name derived from this feature. Each double-legged segment is a resu ...

spider Spiders ( order Araneae) are air-breathing arthropods that have eight legs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species ...

s and

pseudoscorpion Pseudoscorpions, also known as false scorpions or book scorpions, are small, scorpion-like arachnids belonging to the order Pseudoscorpiones, also known as Pseudoscorpionida or Chelonethida. Pseudoscorpions are generally beneficial to humans sin ...

s. Breeding takes place in June and July with females laying clutches of nine to seventeen eggs, usually in chambers hollowed out in rotting logs. The larvae develop inside the eggs, emerging as juveniles in about two months, thus bypassing the free-living larval stage. One or both parents may provide care for the eggs before they hatch.

Status

The clouded salamander is listed as least concern in the IUCN Red List of Threatened Species. This salamander's favored microhabitat of tree stumps and coarse woody decaying debris are being replaced by changes in forestry practices. However, plans to conserve the

spotted owl The spotted owl (''Strix occidentalis'') is a species of true owl. It is a resident species of old-growth forests in western North America, where it nests in tree hollows, old bird of prey nests, or rock crevices. Nests can be between high a ...

(''Strix occidentalis'') and the marbled murrelet (''Brachyramphus marmoratus'') may help to reduce the decline in numbers of clouded salamanders.

References

{{Taxonbar, from=Q1311167 Aneides Amphibians described in 1869 Taxonomy articles created by Polbot