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''Andrias scheuchzeri'' is an
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
species of
giant salamander The Cryptobranchidae are a family of fully aquatic salamanders commonly known as the giant salamanders. They include the largest living amphibians. The family is native to China, Japan, and the eastern United States. They constitute one of two li ...
belonging to the genus ''
Andrias ''Andrias'' is a genus of giant salamanders. It includes the largest salamanders in the world, with ''A. japonicus'' reaching a length of , and ''A. sligoi'' reaching . While extant species are only known from East Asia, several extinct species i ...
'', which also contains the closely related living Asian giant salamanders. It is known from
Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the ...
to
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58Central Europe Central Europe is an area of Europe between Western Europe and Eastern Europe, based on a common historical, social and cultural identity. The Thirty Years' War (1618–1648) between Catholicism and Protestantism significantly shaped the area' ...
, but possibly as far east as
Western Siberia Western Siberia or West Siberia (russian: Западная Сибирь, Zapadnaya Sibir'; kk, Батыс Сібір) is a part of the larger region of Siberia that is mostly located in the Russian Federation. It lies between the Ural region an ...
and eastern
Kazakhstan Kazakhstan, officially the Republic of Kazakhstan, is a transcontinental country located mainly in Central Asia and partly in Eastern Europe. It borders Russia to the north and west, China to the east, Kyrgyzstan to the southeast, Uzbeki ...
.


History

In his book ''Lithographia Helvetica'' from 1726,
Johann Jakob Scheuchzer Johann Jakob Scheuchzer (2 August 1672 – 23 June 1733) was a Swiss scholar born at Zürich. Herbarium deluvianum Zürich, Zwingli-Platz ( Grossmünster) : Former home of Konrad von Mure († 1280) and the house, where Johann Jakob Scheuch ...
described a
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
found in
Öhningen Öhningen is a municipality on the western edge of Lake Constance where it forms the border between Switzerland and the district of Konstanz (or Constance) in Baden-Württemberg in Germany. World heritage site It is home to one or more prehistor ...
as ''Homo diluvii testis'' (
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
: ''Man, witness of the
Deluge A deluge is a large downpour of rain, often a flood. The Deluge refers to the flood narrative in the Biblical book of Genesis. Deluge may also refer to: History *Deluge (history), the Swedish and Russian invasion of the Polish-Lithuanian Comm ...
''), believing it to be the remains of a human that drowned in the
biblical The Bible (from Koine Greek , , 'the books') is a collection of religious texts or scriptures that are held to be sacred in Christianity, Judaism, Samaritanism, and many other religions. The Bible is an anthologya compilation of texts of a ...
Deluge. The fossil was about 1 m (3 ft) long, lacked its tail and hind legs, and could thus be interpreted as showing some resemblance to the remains of a violently trampled human child. In 1758, the first to doubt his theory in print was
Johannes Gessner Johannes Gessner (18 March 1709 – 6 May 1790) was a Swiss mathematician, physicist, botanist, mineralogist and physician. He is seen as the founder of the "Naturforschende Gesellschaft in Zürich". Gessner was born and died in Zürich, whe ...
, who thought it was a giant
catfish Catfish (or catfishes; order Siluriformes or Nematognathi) are a diverse group of ray-finned fish. Named for their prominent barbels, which resemble a cat's whiskers, catfish range in size and behavior from the three largest species alive, ...
(Siluris). In 1787
Petrus Camper Petrus Camper FRS (11 May 1722 – 7 April 1789), was a Dutch physician, anatomist, physiologist, midwife, zoologist, anthropologist, palaeontologist and a naturalist in the Age of Enlightenment. He was one of the first to take an interest in ...
thought it was a lizard (Lacerta); at that time, scholars and the scientific community generally did not differentiate between reptiles and amphibians. In 1802
Martin van Marum Martin(us) van Marum (20 March 1750, Delft – 26 December 1837, Haarlem) was a Dutch physician, inventor, scientist and teacher, who studied medicine and philosophy in Groningen. Van Marum introduced modern chemistry in the Netherlands after ...
bought this fossil for
Teyler's Museum Teylers Museum () is an art, natural history, and science museum in Haarlem, Netherlands. Established in 1778, Teylers Museum was founded as a centre for contemporary art and science. The historic centre of the museum is the neoclassical Oval R ...
in
Haarlem Haarlem (; predecessor of ''Harlem'' in English) is a city and municipality in the Netherlands. It is the capital of the province of North Holland. Haarlem is situated at the northern edge of the Randstad, one of the most populated metropoli ...
from Scheuchzer's grandson in
Zürich Zürich () is the list of cities in Switzerland, largest city in Switzerland and the capital of the canton of Zürich. It is located in north-central Switzerland, at the northwestern tip of Lake Zürich. As of January 2020, the municipality has 43 ...
, along with a fossilized swordfish, for 14
Louis d'or The Louis d'or () is any number of French coins first introduced by Louis XIII in 1640. The name derives from the depiction of the portrait of King Louis on one side of the coin; the French royal coat of arms is on the reverse. The coin was re ...
. It can still be seen in Teyler's Museum, in the original showcase. Seven years later, the fossil again came under scrutiny when the famous
Georges Cuvier Jean Léopold Nicolas Frédéric, Baron Cuvier (; 23 August 1769 – 13 May 1832), known as Georges Cuvier, was a French natural history, naturalist and zoology, zoologist, sometimes referred to as the "founding father of paleontology". Cuvier ...
published an article in which he claimed the fossil was "nothing but a salamander, or rather a
proteus In Greek mythology, Proteus (; Ancient Greek: Πρωτεύς, ''Prōteus'') is an early prophetic sea-god or god of rivers and oceanic bodies of water, one of several deities whom Homer calls the "Old Man of the Sea" ''(hálios gérôn)''. ...
of gigantic dimensions and of an unknown species". He proceeded to examine the fossil in Haarlem, by then a part of the French Empire, in 1811. After hacking away gently at the fossil, he uncovered the foremost limbs and the specimen was recognized as a giant salamander. The difference in color of the stone shows what Scheuchzer saw and what Cuvier later could see. The specimen was renamed ''
Salamandra ''Salamandra'' is a genus of seven species of salamanders localized in central and southern Europe, Northern Africa, and western Asia. List of species References External links Salamandraat Fauna Europaea * Salamandraat Animal Diversity W ...
scheuchzeri'' by Friedrich Holl in 1831. The
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''Andrias'' was only coined six years later by
Johann Jakob von Tschudi Johann Jakob von Tschudi (25 July 1818 – 8 October 1889) was a Switzerland, Swiss Natural history, naturalist, explorer and diplomat. Biography Tschudi was born in Glarus to Johann Jakob Tschudi, a merchant, and Anna Maria Zwicky. He studied ...
. In doing so, both the genus, ''Andrias'' (which means ''image of man''), and the specific name, ''scheuchzeri'', ended up honouring Scheuchzer and his beliefs. The Teylers Museum has several other specimens in their collection in addition to this one.


Description

A number of traits are characteristic of ''Andrias'' ''scheuchzeri'', including strongly curved dentaries, the pars facialis of the skull having a rough inner surface with ridges and depressions, and a number of characters relating to the arrangement and shape of the teeth.


Distribution

In Central Europe, fossil are known from the Late
Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the ...
(
Chattian The Chattian is, in the geologic timescale, the younger of two ages or upper of two stages of the Oligocene Epoch/Series. It spans the time between . The Chattian is preceded by the Rupelian and is followed by the Aquitanian (the lowest stage ...
) to Late
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58Piacenzian The Piacenzian is in the international geologic time scale the upper stage (stratigraphy), stage or latest age (geology), age of the Pliocene. It spans the time between 3.6 ± 0.005 year#SI prefix multipliers, Ma and 2.588 ± 0.005 Ma (million yea ...
) of Germany, the Early
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
of the Czech Republic, the Late Miocene of Austria, and the Middle or Late Miocene of Hungary. Some reports have claimed that ''Andrias scheuchzeri'' was distributed as far east as
Western Siberia Western Siberia or West Siberia (russian: Западная Сибирь, Zapadnaya Sibir'; kk, Батыс Сібір) is a part of the larger region of Siberia that is mostly located in the Russian Federation. It lies between the Ural region an ...
and the Zaissan Basin on the border between China and Kazakhstan.


Ecology

''Andrias scheuchzeri'' inhabited ponds, lakes and rivers, unlike modern Asian giant salamanders, which prefer clear, nutrient poor (
oligotrophic An oligotroph is an organism that can live in an environment that offers very low levels of nutrients. They may be contrasted with copiotrophs, which prefer nutritionally rich environments. Oligotrophs are characterized by slow growth, low rates of ...
) rivers and streams.


In fiction

The fictional descendants of ''Andrias scheuchzeri'' are the primary antagonists in Karel Čapek's 1936
science fiction Science fiction (sometimes shortened to Sci-Fi or SF) is a genre of speculative fiction which typically deals with imaginative and futuristic concepts such as advanced science and technology, space exploration, time travel, parallel unive ...
novel ''
War with the Newts ''War with the Newts'' (''Válka s Mloky'' in the original Czech), also translated as ''Salamander Wars'', is a 1936 satirical science fiction novel by Czech author Karel Čapek. It concerns the discovery in the Pacific of a sea-dwelling race, ...
''.


Sources


Bibliography

* Richard Ellis (2001), ''Aquagenesis: The Origin and Evolution of Life in the Sea'' (London: Penguin). * Robert Hofrichter (2000), ''Amphibians: The World of Frogs, Toads, Salamanders and Newts'' (Toronto: Key Porter Books) * J. Alan Holman (2006), ''Fossil Salamanders of North America (Life of the Past)'' (Bloomington: Indiana University Press) * J. William Schopf (2001), ''Cradle of Life: The Discovery of Earth's Earliest Fossils'' (Princeton: Princeton University Press) {{DEFAULTSORT:Andrias Scheuchzeri Cryptobranchidae Paleogene animals of Europe Miocene amphibians Cenozoic salamanders Fossil taxa described in 1831 Oligocene species first appearances Pliocene species extinctions