Andean Emerald
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The Andean emerald (''Uranomitra franciae'') is a species of
hummingbird Hummingbirds are birds native to the Americas and comprise the biological family Trochilidae. With about 361 species and 113 genera, they occur from Alaska to Tierra del Fuego, but the vast majority of the species are found in the tropics aro ...
in the "emeralds", tribe Trochilini of subfamily Trochilinae. It is found in
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
,
Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ''Eku ...
, and
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = Seal (emblem), National seal , national_motto = "Fi ...
.


Taxonomy and systematics

The Andean emerald was formerly placed in the genus '' Amazilia''. A
molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...
study published in 2014 found that the genus ''Amazilia'' was polyphyletic. In the revised classification to create
monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ...
genera, the Andean emerald was moved by most taxonomic systems to the resurrected genus ''Uranomitra'' that had been introduced in 1854 by
Ludwig Reichenbach Heinrich Gottlieb Ludwig Reichenbach (8 January 1793 – 17 March 1879) was a German botanist and ornithologist. It was he who first requested Leopold Blaschka to make a set of glass marine invertebrate models for scientific education and museu ...
.Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, A. Jaramillo, D. F. Lane, J. F. Pacheco, M. B. Robbins, F. G. Stiles, and K. J. Zimmer. Version 24 July 2022. A classification of the bird species of South America. American Ornithological Society. https://www.museum.lsu.edu/~Remsen/SACCBaseline.htm retrieved July 24, 2022Clements, J. F., T. S. Schulenberg, M. J. Iliff, S. M. Billerman, T. A. Fredericks, J. A. Gerbracht, D. Lepage, B. L. Sullivan, and C. L. Wood. 2021. The eBird/Clements checklist of Birds of the World: v2021. Downloaded from https://www.birds.cornell.edu/clementschecklist/download/ Retrieved August 25, 2021 However,
BirdLife International BirdLife International is a global partnership of non-governmental organizations that strives to conserve birds and their habitats. BirdLife International's priorities include preventing extinction of bird species, identifying and safeguarding ...
's ''
Handbook of the Birds of the World The ''Handbook of the Birds of the World'' (HBW) is a multi-volume series produced by the Spanish publishing house Lynx Edicions in partnership with BirdLife International. It is the first handbook to cover every known living species of bird. T ...
'' (HBW) retains it in ''Amazilia''. The Andean emerald is the only species in genus ''Uranomitra''. These three
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are recognized: * ''U. f. franciae'' (Bourcier & Mulsant, 1846) * ''U. f. viridiceps'' (Gould, 1860) * ''U. f. cyanocollis'' (Gould, 1853) Andean emerald (Uranomitra franciae franciae) male Cundinamarca.jpg, male ''U. f. franciae''. Colombia Andean emerald (Uranomitra franciae franciae) male in flight Las Tangaras.jpg, male ''U. f. franciae''. Colombia TAN 0096 - Andean Emerald.jpg, male ''U. f. viridiceps'',
Bellavista Cloud Forest Reserve The Bellavista Cloud Forest Reserve is a certified conservation area on the North-Western slopes of the Andean mountain range and is located 52 km from the Ecuadorian capital city of Quito. The lodge itself comprises guest rooms, 10 km ...
,
Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ''Eku ...


Description

The Andean emerald is long. Males weigh about and females about . Both sexes of all subspecies have a straight to slightly decurved bill of medium length with a blackish
maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The t ...
and a coral red
mandible In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower tooth, teeth in place. The mandible sits beneath the maxilla. It is the only movabl ...
with a dark tip. Adult males of the
nominate subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
''U. f. franciae'' have a glittering violet-blue crown and glittering golden- to emerald green cheeks and neck. Their upperparts and flanks are light green to golden-green with a coppery tinge to the uppertail
coverts A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which, as the name implies, cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are sm ...
. The center of their underparts is white. Their tail is bronze-green to copperish. The adult female is similar to the male but its crown is turquoise-blue to greenish and less glittering. Juveniles are similar to the adult female and also have brownish edges on the upperparts' feathers and grayish brown flanks.Weller, A.A., P. F. D. Boesman, and G. M. Kirwan (2021). Andean Emerald (''Uranomitra franciae''), version 1.1. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.andeme1.01.1 retrieved September 10, 2022 Subspecies ''U. f. viridiceps'' differs from the nominate with a shorter tail and a green crown. ''U. f. cyanocollis'' also differs slightly from the nominate; the blue of its crown extends to the nape.


Distribution and habitat

The subspecies of Andean emerald are found thus: * ''U. f. franciae'', Andes of northwestern and central Colombia * ''U. f. viridiceps'', Andes from southwestern Colombia's Nariño Department through western Ecuador to
Loja Province Loja Province () is one of 24 provinces in Ecuador and shares its southern border on the west with El Oro Province, on the north with El Azuay, and on the east with Zamora-Chinchipe. Founded on its present site in 1548 by Captain Alonso de Mer ...
* ''U. f. cyanocollis'', Andes of southeastern Ecuador's Zamora-Chinchipe Province and northern Peru's Marañón River valley south and east to La Libertad Province The Andean emerald inhabits
secondary forest A secondary forest (or second-growth forest) is a forest or woodland area which has re-grown after a timber harvest or clearing for agriculture, until a long enough period has passed so that the effects of the disturbance are no longer evident. ...
and the edges and clearings of wet primary forest. In Peru it also occurs in drier shrublands. In elevation it ranges between and is most common above .


Behavior


Movement

The Andean emerald makes seasonal elevational movements that however are not well defined.


Feeding

The Andean emerald forages for nectar at a variety of flowering herbs, vines, and trees, usually from the middle to upper strata of the forest. It forages by
trap-lining In ethology and behavioral ecology, trap-lining or traplining is a feeding strategy in which an individual visits food sources on a regular, repeatable sequence, much as trappers check their lines of traps. Traplining is usually seen in species ...
, visiting a circuit of nectar sources. It sometimes mixes with other hummingbirds at flower patches but is subordinate to territorial species. In addition to nectar it feeds on insects by
hawking Hawking may refer to: People * Stephen Hawking (1942–2018), English theoretical physicist and cosmologist *Hawking (surname), a family name (including a list of other persons with the name) Film * ''Hawking'' (2004 film), about Stephen Haw ...
from a perch.


Breeding

Andean emerald breeding activity has been documented at almost every time of year. It builds a cup nest of treefern scales and plant fibers bound with spiderweb and with lichen on the outside. It typically sites it about above the ground. The female incubates the clutch of two eggs for 19 to 24 days and fledging occurs about 16 to 22 days after hatch.


Vocalization

The Andean emerald has a quite variable song, typically described as "a repeated complex phrase of high-pitched squeaky whistles, trills and scratchy notes." It also makes "a high-pitched 'tsip'" call, and during social interactions "scratchy squeaky chatters".


Status

The
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
has assessed the Andean Emerald as being of Least Concern, though its population size and trend are unknown. No immeditate threats have been identified. It is considered uncommon to fairly common and is found in at least one protected area.


References

{{Taxonbar, from=Q760693 Andean emerald Birds of the Andes Birds of the Northern Andes Hummingbird species of South America Andean emerald Andean emerald Andean emerald Taxobox binomials not recognized by IUCN