Amplirhagada Berthierana
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''Amplirhagada'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of air-breathing
land snail A land snail is any of the numerous species of snail that live on land, as opposed to the sea snails and freshwater snails. ''Land snail'' is the common name for terrestrial gastropod mollusks that have shells (those without shells are known as ...
s,
terrestrial Terrestrial refers to things related to land or the planet Earth. Terrestrial may also refer to: * Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on or near the ground, as opposed to ...
pulmonate Pulmonata or pulmonates, is an informal group (previously an order, and before that a subclass) of snails and slugs characterized by the ability to breathe air, by virtue of having a pallial lung instead of a gill, or gills. The group includ ...
gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. T ...
mollusk Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is e ...
s in the family
Camaenidae Camaenidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Helicoidea, the typical snails and their allies. This is one of the most diverse families in the clade Stylommatophora. These ...
. This genus is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to the
Kimberley Kimberly or Kimberley may refer to: Places and historical events Australia * Kimberley (Western Australia) ** Roman Catholic Diocese of Kimberley * Kimberley Warm Springs, Tasmania * Kimberley, Tasmania a small town * County of Kimberley, a ...
region in northwesternmost
Western Australia Western Australia (commonly abbreviated as WA) is a state of Australia occupying the western percent of the land area of Australia excluding external territories. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to th ...
, where it represents the most species-rich
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of land snails.Köhler, F. (2010). Uncovering local endemism in the Kimberley, Western Australia: Description of new species of the genus ''Amplirhagada'' Iredale, 1933 (Pulmonata, Camaenidae). ''Records of the Australian Museum'' 62(2): 217-284.


Characteristic features

''Amplirhagada''-species exhibit variable shells ranging from broadly conical to highly turreted in shape. Shells are comparatively large, ranging from about 10 to 25 mm in height and from about 15 to 30 mm in diameter. Shells are often banded but uniformly brown species are also known. The umbilicus is narrowly winding and open, partly or completely concealed by the
columella Lucius Junius Moderatus Columella (; Arabic: , 4 – ) was a prominent writer on agriculture in the Roman Empire. His ' in twelve volumes has been completely preserved and forms an important source on Roman agriculture, together with the wo ...
r reflection. Typical features of the genital organs comprise presence of
penial sheath In male human anatomy, the foreskin, also known as the prepuce, is the double-layered fold of skin, mucosal and muscular tissue at the distal end of the human penis that covers the glans and the urinary meatus. The foreskin is attached to the g ...
and penial verge and absence of an
epiphallus The reproductive system of gastropods (slugs and snails) varies greatly from one group to another within this very large and diverse taxonomic class of animals. Their reproductive strategies also vary greatly, see Mating of gastropods. In many mar ...
.


Species

Currently, 56 species are known.Solem, A. 1981. Camaenid land snails from Western and central Australia. II. Taxa from the Kimberley, ''Amplirhagada'' Iredale, 1933. ''Records of the Western Australian Museum'', Supplement 11: 147-320. These include: * '' Amplirhagada alta'' Solem, 1981 * '' Amplirhagada anderdonensis'' Köhler, 2010 * '' Amplirhagada astuta'' ( Iredale, 1939) * '' Amplirhagada basilica'' Köhler, 2010 * '' Amplirhagada berthierana'' Köhler, 2010 * ''
Amplirhagada boongareensis ''Amplirhagada'' is a genus of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Camaenidae. This genus is endemic to the Kimberley region in northwesternmost Western Australia, where it represents the most specie ...
'' Köhler, 2010 * ''
Amplirhagada buffonensis ''Amplirhagada'' is a genus of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Camaenidae. This genus is endemic to the Kimberley region in northwesternmost Western Australia, where it represents the most specie ...
'' Köhler, 2010 * '' Amplirhagada burnerensis'' ( E.A. Smith, 1894) * '' Amplirhagada burrowsena'' Iredale, 1939 * '' Amplirhagada cambridgensi''s Solem, 1988 * '' Amplirhagada camdenensis'' Köhler, 2010 * '' Amplirhagada carinata'' Solem, 1981 * '' Amplirhagada castra'' Solem, 1981 * '' Amplirhagada combeana'' Iredale, 1938 * '' Amplirhagada confusa'' Solem, 1981 * '' Amplirhagada crystalla'' ( Solem, 1981) * '' Amplirhagada decora'' Köhler, 2010 * '' Amplirhagada depressa'' ( Solem, 1981) * '' Amplirhagada descartesana'' Köhler, 2010 * '' Amplirhagada drysdaleana'' Solem, 1981 * '' Amplirhagada dubitabile'' Köhler, 2010 * '' Amplirhagada elevata'' Solem, 1981 * '' Amplirhagada euroa'' Köhler, 2010 * '' Amplirhagada gemina'' Köhler, 2010 * '' Amplirhagada gibsoni'' Köhler, 2010 * '' Amplirhagada herbertena'' Iredale, 1939 * '' Amplirhagada imitata'' ( E.A. Smith, 1894) * '' Amplirhagada indistincta'' Köhler, 2010 * '' Amplirhagada intermedia'' ( Solem, 1981) * '' Amplirhagada kalumburuana'' Solem, 1981 * '' Amplirhagada katerana'' Solem, 1981 * '' Amplirhagada kessneri'' Köhler, 2010 * '' Amplirhagada kimberleyana'' Köhler, 2010 * '' Amplirhagada lamarckiana'' Köhler, 2010 * '' Amplirhagada mckenziei'' Köhler, 2010 * '' Amplirhagada mitchelliana'' Solem, 1981 * '' Amplirhagada montalevitensis'' ( E.A. Smith, 1894) * '' Amplirhagada montesqieuana'' Köhler, 2010 * '' Amplirhagada napierana'' Solem, 1981 * '' Amplirhagada novelta'' Iredale, 1939 * '' Amplirhagada osmondi'' Solem, 1988 * '' Amplirhagada percita'' ( Iredale, 1939) * '' Amplirhagada ponderi'' Köhler, 2010 * '' Amplirhagada puescheli'' Köhler, 2010 * '' Amplirhagada pusilla'' Solem, 1981 * '' Amplirhagada questroana'' Solem, 1981 * '' Amplirhagada regia'' Köhler, 2010 * '' Amplirhagada solemiana'' Köhler, 2010 * '' Amplirhagada sphaeroidea'' Köhler, 2010 * '' Amplirhagada sykesi'' ( E.A. Smith, 1894) * '' Amplirhagada tricenaria'' Köhler, 2010 * '' Amplirhagada uwinsensis'' Köhler, 2010 * '' Amplirhagada varia'' Solem, 1981 * '' Amplirhagada wilsoni'' Solem, 1981 * '' Amplirhagada yorkensis'' Köhler, 2010


References

{{Taxonbar, from=Q3079941 Camaenidae Taxonomy articles created by Polbot