The copper underwing, humped green fruitworm or pyramidal green fruitworm (''Amphipyra pyramidea'') is a

moth Moths are a group of insects that includes all members of the order Lepidoptera that are not Butterfly, butterflies. They were previously classified as suborder Heterocera, but the group is Paraphyly, paraphyletic with respect to butterflies (s ...

of the family

Noctuidae The Noctuidae, commonly known as owlet moths, cutworms or armyworms, are a family (biology), family of moths. Taxonomically, they are considered the most controversial family in the superfamily Noctuoidea because many of the clades are constantly ...

. The species was first described by

Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...

in his 1758 10th edition of ''Systema Naturae''.

Distribution

This species can be found across the

Palaearctic The Palearctic or Palaearctic is a biogeographic realm of the Earth, the largest of eight. Confined almost entirely to the Eastern Hemisphere, it stretches across Europe and Asia, north of the foothills of the Himalayas, and North Africa. Th ...

region including Europe,

North Africa North Africa (sometimes Northern Africa) is a region encompassing the northern portion of the African continent. There is no singularly accepted scope for the region. However, it is sometimes defined as stretching from the Atlantic shores of t ...

, the

Near East The Near East () is a transcontinental region around the Eastern Mediterranean encompassing the historical Fertile Crescent, the Levant, Anatolia, Egypt, Mesopotamia, and coastal areas of the Arabian Peninsula. The term was invented in the 20th ...

Iran Iran, officially the Islamic Republic of Iran (IRI) and also known as Persia, is a country in West Asia. It borders Iraq to the west, Turkey, Azerbaijan, and Armenia to the northwest, the Caspian Sea to the north, Turkmenistan to the nort ...

, southern

Siberia Siberia ( ; , ) is an extensive geographical region comprising all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has formed a part of the sovereign territory of Russia and its predecessor states ...

, northern

India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...

Korea Korea is a peninsular region in East Asia consisting of the Korean Peninsula, Jeju Island, and smaller islands. Since the end of World War II in 1945, it has been politically Division of Korea, divided at or near the 38th parallel north, 3 ...

and

Japan Japan is an island country in East Asia. Located in the Pacific Ocean off the northeast coast of the Asia, Asian mainland, it is bordered on the west by the Sea of Japan and extends from the Sea of Okhotsk in the north to the East China Sea ...

. It is rather common over the southern half of Britain.

Description

This species has a

wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the opposite wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingsp ...

of 47–54 mm, the female usually slightly larger than the male. The forewings are brown marked with paler

fascia A fascia (; : fasciae or fascias; adjective fascial; ) is a generic term for macroscopic membranous bodily structures. Fasciae are classified as superficial, visceral or deep, and further designated according to their anatomical location. ...

and a pale, dark-centred stigma. The hindwings are a rich bright copper colour (hence the common name of "copper underwing"). It is very similar to

Svensson's copper underwing Svensson's copper underwing (''Amphipyra berbera'') is a moth of the family Noctuidae. The species was first described by Charles E. Rungs in 1949. It is distributed throughout Europe including Russia east to the Urals. This species has a wings ...

(''Amphipyra berbera'') but identification is usually fairly straightforward by looking at some markings on the forewings, the detail of the labial palps and at the underside of the hindwings. This species has a pale area in the centre of the hindwings, contrasting with much darker marginal areas, while in ''A. berbera'' the whole underwing is more or less uniform in colour. See Townsend et al. The

larva A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect development such as insects, some arachnids, amphibians, or cnidarians typically have a larval phase ...

is green with white markings and a pointed hump at the rear end.

Differentiation of ''A. pyramidea'' from ''A. berbera''

File:Amphipyra pyramidea - differenciation from Amphipyra berbera 01.jpg, ''A. pyramidea'' - red arrow points out a difference File:Amphipyra pyramidea - differenciation from Amphipyra berbera 02.jpg, ''A. berbera'' - red arrows point out a difference File:Amphipyra pyramidea - berbera larvae.jpg, Caterpillars of ''A. pyramidea'' (top) and ''A. berbera''

Biology

It is a

univoltine Voltinism is a term used in biology to indicate the number of broods or generations of an organism in a year. The term is most often applied to insects, and is particularly in use in sericulture, where silkworm varieties vary in their voltinism. ...

species. Hatching season is as early as June in some climates but they usually hatch July to October. Eggs are laid in deciduous trees. Larva can be found as early as April in some climates but usually emerge in May–June. ''A pyramidea'' flies at night from August to October and is attracted to light and strongly to sugar. It feeds on a variety of

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...

s and

shrub A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...

s, including ash tree (''

Fraxinus ''Fraxinus'' (), commonly called ash, is a genus of plants in the olive and lilac family, Oleaceae, and comprises 45–65 species of usually medium-to-large trees, most of which are deciduous trees, although some Subtropics, subtropical specie ...

''), privet (''

Ligustrum A privet is a flowering plant in the genus ''Ligustrum''. The genus contains about 50 species of erect, deciduous or evergreen shrubs or trees, with a native distribution from Europe to tropical and subtropical Asia, and with one species each ...

''), honeysuckle (''

Lonicera Honeysuckles are arching shrubs or twining vines in the genus ''Lonicera'' () of the family Caprifoliaceae. The genus includes 158 species native to northern latitudes in North America, Eurasia, and North Africa. Widely known species include ...

''), apple (''

Malus ''Malus'' ( or ) is a genus of about 32–57 species of small deciduous trees or shrubs in the family Rosaceae, including the domesticated orchard apple, crab apples (sometimes known in North America as crabapples) and wild apples. The genus i ...

''), oak (''

Quercus An oak is a hardwood tree or shrub in the genus ''Quercus'' of the beech family. They have spirally arranged leaves, often with lobed edges, and a nut called an acorn, borne within a cup. The genus is widely distributed in the Northern Hemisp ...

''), ''

Rhododendron ''Rhododendron'' (; : ''rhododendra'') is a very large genus of about 1,024 species of woody plants in the Ericaceae, heath family (Ericaceae). They can be either evergreen or deciduous. Most species are native to eastern Asia and the Himalayan ...

'',

rose A rose is either a woody perennial plant, perennial flowering plant of the genus ''Rosa'' (), in the family Rosaceae (), or the flower it bears. There are over three hundred Rose species, species and Garden roses, tens of thousands of cultivar ...

, wild service tree (''

Sorbus ''Sorbus'' is a genus of over 100 species of trees and shrubs in the rose family, Rosaceae. Species of ''Sorbus'' ('' s.str.'') are commonly known as rowan or mountain-ash. The genus used to include species commonly known as whitebeam, cheque ...

'') and lilac (''

Syringa ''Syringa'' is a genus of 12 currently recognized species of flowering plant, flowering woody plants in the olive family or Oleaceae called lilacs. These lilacs are native to woodland and scrub from southeastern Europe to eastern Asia, and wid ...

'').Mazzei, Paolo; Morel, Daniel & Panfili, Raniero
''Moths and Butterflies of Europe and North Africa''
/ref> The species overwinters as an

egg An egg is an organic vessel grown by an animal to carry a possibly fertilized egg cell (a zygote) and to incubate from it an embryo within the egg until the embryo has become an animal fetus that can survive on its own, at which point the ...

. # ''The flight season refers to the

British Isles The British Isles are an archipelago in the Atlantic Ocean, North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner Hebrides, Inner and Outer Hebr ...

. This may vary in other parts of the range.''

Gallery

File: Amphipyra pyramidea larva.jpg, Larva File: Noctuidae - Amphipyra pyramidea.JPG, Moth File:Amphipyra pyramidea.jpg, Mounted specimen File: F Nemos OBA Amphipyra pyramidea.jpg, Illustration

References

* *

External links

''Butterflies & Moths of Switzerland''''Lepiforum e. V.''
Includes photo of genitalia.

{{Taxonbar, from=Q735875 Amphipyrinae Moths of North America Moths of Europe Moths of Asia Moths of Africa Moths described in 1758 Taxa named by Carl Linnaeus