Amphiprion Pacificus
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''Amphiprion pacificus'', is a species of
anemonefish Clownfish or anemonefish are fishes from the subfamily Amphiprioninae in the family Pomacentridae. Thirty species of clownfish are recognized: one in the genus '' Premnas'', while the remaining are in the genus '' Amphiprion''. In the wild, t ...
that is found in the western
Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continen ...
. Like all anemonefishes it forms a
symbiotic Symbiosis (from Greek , , "living together", from , , "together", and , bíōsis, "living") is any type of a close and long-term biological interaction between two different biological organisms, be it mutualistic, commensalistic, or parasit ...
mutualism with
sea anemone Sea anemones are a group of predation, predatory marine invertebrates of the order (biology), order Actiniaria. Because of their colourful appearance, they are named after the ''Anemone'', a terrestrial flowering plant. Sea anemones are classifi ...
s and is unaffected by the stinging tentacles of the host anemone. It is a
sequential hermaphrodite Sequential hermaphroditism (called dichogamy in botany) is a type of hermaphroditism that occurs in many fish, gastropods, and plants. Sequential hermaphroditism occurs when the individual changes its sex at some point in its life. In particular, ...
with a strict sized based dominance hierarchy: the female is largest, the breeding male is second largest, and the male non-breeders get progressively smaller as the hierarchy descends. They exhibit
protandry Sequential hermaphroditism (called dichogamy in botany) is a type of hermaphroditism that occurs in many fish, gastropods, and plants. Sequential hermaphroditism occurs when the individual changes its sex at some point in its life. In particular, ...
, meaning the breeding male will change to female if the sole breeding female dies, with the largest non-breeder becomes the breeding male. The fish's natural diet includes
zooplankton Zooplankton are the animal component of the planktonic community ("zoo" comes from the Greek word for ''animal''). Plankton are aquatic organisms that are unable to swim effectively against currents, and consequently drift or are carried along by ...
.


Description

The body of ''A. pacificus'' is pinkish brown dark brown, usually grading to yellowish or orange on lower side of body including abdomen. It is a member of the skunk complex with the characteristic white stripe along the dorsal ridge line, from the midline of the snout through to the base of
caudal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
. The fins are whitish to semi-translucent. It has 9 dorsal spines, 2 anal spines, 18-20 dorsal soft rays and 12-13 anal soft rays. The four specimens that were the basis of the description ranged from .


Color variations

Some anemonefish species have color variations based on geographic location, sex and host anemone. ''A. pacificus'', like other members of the skunk complex does not show any of these variations.


Similar species

'' A. akallopisos'' is almost identical, however there is a wide geographic separation such that geographic location is the easiest distinction. Despite the similarity in appearance, genetic results indicate that ''A. pacificus'' is well differentiated from ''A. akallopisos'' and is more closely related to '' A. sandaracinos'' which has a uniform coloration and the white stripe on the dorsal ridge extends onto the superior lip. 3 other species of anemonefish are found within its range, ''A. barberi'', ''A. chrysopterus'' and ''A. clarkii'', however these are easily distinguished by the absence of the white stripe on the dorsal ridge. Image:Amphiprion akallopisos 2010.JPG, ''A. akallopisos'' (Skunk anemonefish) Image:Amphiprion sandaracinos.jpg, ''A. sandaracinos'' (Orange anemonefish) showing the broader white stripe extending to the upper lip.


Distribution and Habitat

''A. pacificus'' is found the western Pacific at
Wallis Island Wallis (Wallisian: ''Uvea'') is a Polynesian atoll/island in the Pacific Ocean belonging to the French overseas collectivity (''collectivité d'outre-mer'', or ''COM'') of Wallis and Futuna. It lies north of Tonga, northeast of Fiji, east-nort ...
,
Tonga Tonga (, ; ), officially the Kingdom of Tonga ( to, Puleʻanga Fakatuʻi ʻo Tonga), is a Polynesian country and archipelago. The country has 171 islands – of which 45 are inhabited. Its total surface area is about , scattered over in ...
,
Fiji Fiji ( , ,; fj, Viti, ; Fiji Hindi: फ़िजी, ''Fijī''), officially the Republic of Fiji, is an island country in Melanesia, part of Oceania in the South Pacific Ocean. It lies about north-northeast of New Zealand. Fiji consists ...
and Samoan Islands, a regional hotspot of endemism. It is not common within its range and is generally found at depths of .


Host anemones

The relationship between anemonefish and their host sea anemones is not random and instead is highly
nested ''Nested'' is the seventh studio album by Bronx-born singer, songwriter and pianist Laura Nyro, released in 1978 on Columbia Records. Following on from her extensive tour to promote 1976's ''Smile'', which resulted in the 1977 live album ''Season ...
in structure. ''A. pacificus'' is highly specialised, being hosted by only 1 out of the 6 host anemones found in the region. ''A. pacificus'' is hosted by the following species of anemone: *''
Heteractis magnifica ''Heteractis magnifica'', also known by the common names magnificent sea anemone or Ritteri anemone, is a species of sea anemone belonging to the Stichodactylidae family native to the Indo-Pacific area. Description The magnificent sea anemone is ...
'' magnificent sea anemone


Conservation status

Anemonefish and their host anemones are found on coral reefs and face similar environmental issues. Like
coral Corals are marine invertebrates within the class Anthozoa of the phylum Cnidaria. They typically form compact colonies of many identical individual polyps. Coral species include the important reef builders that inhabit tropical oceans and sec ...
s, anemone's contain intracellular endosymbionts, zooxanthellae, and can suffer from
bleaching Bleach is the generic name for any chemical product that is used industrially or domestically to remove color (whitening) from a fabric or fiber or to clean or to remove stains in a process called bleaching. It often refers specifically, to ...
due to triggers such as increased water temperature or
acidification Acidification may refer to: * Ocean acidification, decrease in the pH of the Earth's oceans * Freshwater acidification, atmospheric depositions and soil leaching of SOx and NOx * Soil acidification, buildup of hydrogen cations, which reduces the ...
. Characteristics known to elevate the risk of extinction are small geographic range, small local population and extreme habitat specialisation. ''A. pacificus'' has each of these characteristics, being an endemic species, with small local population and only one host anemone suggest that it is of particular conservation concern. This species was not evaluated in the 2012 release of the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biol ...
.


References


External links

* * {{Taxonbar, from=Q219440
pacificus Pacificus was a disciple of St. Francis of Assisi, born probably near Ascoli, Italy, in the second half of the twelfth century; died, it is thought, at Lens, France, around 1234. Poet Laureate Local authors identify him with a certain William of ...
Fish described in 2010