Amphiprion Melanopus
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''Amphiprion melanopus'', also known as the cinnamon clownfish, fire clownfish, red and black anemonefish, black-backed anemonefish or dusky anemonefish is a widely distributed
anemonefish Clownfish or anemonefish are fishes from the subfamily Amphiprioninae in the family Pomacentridae. Thirty species of clownfish are recognized: one in the genus '' Premnas'', while the remaining are in the genus '' Amphiprion''. In the wild, t ...
chiefly found in the western and southern parts of the
Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the contin ...
.. The species scientific name 'melanopus' is
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
, meaning black feet in reference to the black pelvic fins. Like all anemonefishes it forms a symbiotic mutualism with sea anemones and is unaffected by the stinging tentacles of the host anemone. It is a
sequential hermaphrodite Sequential hermaphroditism (called dichogamy in botany) is a type of hermaphroditism that occurs in many fish, gastropods, and plants. Sequential hermaphroditism occurs when the individual changes its sex at some point in its life. In particular, ...
with a strict sized based
dominance hierarchy In biology, a dominance hierarchy (formerly and colloquially called a pecking order) is a type of social hierarchy that arises when members of animal social groups interact, creating a ranking system. A dominant higher-ranking individual is so ...
: the female is largest, the breeding male is second largest, and the male non-breeders get progressively smaller as the hierarchy descends. They exhibit
protandry Sequential hermaphroditism (called dichogamy in botany) is a type of hermaphroditism that occurs in many fish, gastropods, and plants. Sequential hermaphroditism occurs when the individual changes its sex at some point in its life. In particular, ...
, meaning the breeding male will change to female if the sole breeding female dies, with the largest non-breeder becomes the breeding male.


Description

''A. melanopus'' adults can grow to 12 cm (4.7 inches). The body of ''A. melanopus'' is a dark red to orange with mahogany sides. Juveniles and adults have a white head band, which is wide and starts behind the eye, which may gave a blue tint. The
dorsal Dorsal (from Latin ''dorsum'' ‘back’) may refer to: * Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism * Dorsal, positioned on top of an aircraft's fuselage * Dorsal c ...
and
caudal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as ...
s of the fish have a lighter color than the rest of the fish and can sometimes be a cinnamon color. The
pelvic The pelvis (plural pelves or pelvises) is the lower part of the trunk, between the abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also called bony pelvis, or pelvic skeleton). The p ...
and anal fins are usually black.


Similar species

Historically anemonefish have been identified by morphological features, color pattern in the field, while in a laboratory other features such as scalation of the head, tooth shape and body proportions. These features have been used to group species into 6 complexes. ''A. melanopus'' is in the tomato complex and so is similar to other members of the complex. '' A. rubrocinctus'' and ''A. melanopus'' are similar in appearance but geographically distinct with A. rubrocinctus limited to north western Australia. ''A. barberi'' was previously thought to be a geographic variation of ''A. melanopus'' and there are significant color-pattern differences, with A. melanopus having a dark brown or blackish body, compared to the red orange of A. barberi. ''A. melanopus'' also has 19-26 spines in the upper-opercular series while A. barberi has only 11–19. Image:Amphiprion melanopus RLS.jpg, ''A. melanopus'' (Red & Black anemonefish) Image:Anemonefishfiji.jpg, '' A. barberi'' (Barber's anemonefish) Image:Amphiprion rubrocinctus RLS.jpg, '' A. rubrocinctus'' (Australian anemonefish)


Distribution and habitat

''A. melanopus'' is widely distributed in the western Pacific, from the
Great Barrier Reef The Great Barrier Reef is the world's largest coral reef system composed of over 2,900 individual reefs and 900 islands stretching for over over an area of approximately . The reef is located in the Coral Sea, off the coast of Queensland, ...
north to the
Marshall Islands The Marshall Islands ( mh, Ṃajeḷ), officially the Republic of the Marshall Islands ( mh, Aolepān Aorōkin Ṃajeḷ),'' () is an independent island country and microstate near the Equator in the Pacific Ocean, slightly west of the Intern ...
and
Guam Guam (; ch, Guåhan ) is an organized, unincorporated territory of the United States in the Micronesia subregion of the western Pacific Ocean. It is the westernmost point and territory of the United States (reckoned from the geographic cent ...
,
New Guinea New Guinea (; Hiri Motu: ''Niu Gini''; id, Papua, or , historically ) is the world's second-largest island with an area of . Located in Oceania in the southwestern Pacific Ocean, the island is separated from Australia by the wide Torr ...
and from
Vanuatu Vanuatu ( or ; ), officially the Republic of Vanuatu (french: link=no, République de Vanuatu; bi, Ripablik blong Vanuatu), is an island country located in the South Pacific Ocean. The archipelago, which is of volcanic origin, is east of no ...
and New Caledonia to eastern
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
. It was previously thought to be present in Fiji,
Tonga Tonga (, ; ), officially the Kingdom of Tonga ( to, Puleʻanga Fakatuʻi ʻo Tonga), is a Polynesian country and archipelago. The country has 171 islands – of which 45 are inhabited. Its total surface area is about , scattered over in ...
, and the
Samoan Islands The Samoan Islands ( sm, Motu o Sāmoa) are an archipelago covering in the central South Pacific, forming part of Polynesia and of the wider region of Oceania. Administratively, the archipelago comprises all of the Independent State of Samoa an ...
, however that fish has now been described as a separate species, ''A. barberi''.


Host anemones

The relationship between anemonefish and their host sea anemones is not random and instead is highly
nested ''Nested'' is the seventh studio album by Bronx-born singer, songwriter and pianist Laura Nyro, released in 1978 on Columbia Records. Following on from her extensive tour to promote 1976's ''Smile'', which resulted in the 1977 live album '' Seas ...
in structure. ''A. melanopus'' is a generalist, being usually hosted by 1 anemone, but also hosted by 2 other anemones. It is hosted by the following 3 out of the 10 host anemones: * '' Entacmaea quadricolor'' - bubble-tip or purple base anemone (usually) * '' Heteractis crispa'' - leathery sea anemone (occasionally) * '' Heteractis magnifica'' - magnificent sea anemone (rarely) Image:Amphiprion melanopus (Red and black anemonefish) in Entacmaea quadricolor (Bubble sea anemones).jpg, ''A. melanopus'' '' Entacmaea quadricolor'' (Bubble tip anemone) Image:Amphiprion melanopus (Black anemonefish) in Heteractis magnifica (Magnificent anemone).jpg, ''A. melanopus'' in '' Heteractis magnifica'' - magnificent sea anemone


Diet

Like all anemonefish, ''A. melanopus'' is
omnivorous An omnivore () is an animal that has the ability to eat and survive on both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize the nut ...
and its diet is based on
zooplankton Zooplankton are the animal component of the planktonic community ("zoo" comes from the Greek word for ''animal''). Plankton are aquatic organisms that are unable to swim effectively against currents, and consequently drift or are carried along by ...
, small benthic
crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group can ...
s and algaes.


Conservation status

Anemonefish and their host anemones are found on coral reefs and face similar
environmental issues Environmental issues are effects of human activity on the biophysical environment, most often of which are harmful effects that cause environmental degradation. Environmental protection is the practice of protecting the natural environment on t ...
. Like
coral Corals are marine invertebrates within the class Anthozoa of the phylum Cnidaria. They typically form compact colonies of many identical individual polyps. Coral species include the important reef builders that inhabit tropical oceans and ...
s, anemone's contain intracellular
endosymbiont An ''endosymbiont'' or ''endobiont'' is any organism that lives within the body or cells of another organism most often, though not always, in a mutualistic relationship. (The term endosymbiosis is from the Greek: ἔνδον ''endon'' "within ...
s,
zooxanthellae Zooxanthellae is a colloquial term for single-celled dinoflagellates that are able to live in symbiosis with diverse marine invertebrates including demosponges, corals, jellyfish, and nudibranchs. Most known zooxanthellae are in the genus ''S ...
, and can suffer from
bleaching Bleach is the generic name for any chemical product that is used industrially or domestically to remove color (whitening) from a fabric or fiber or to clean or to remove stains in a process called bleaching. It often refers specifically, to ...
due to triggers such as increased water temperature or
acidification Acidification may refer to: * Ocean acidification, decrease in the pH of the Earth's oceans * Freshwater acidification, atmospheric depositions and soil leaching of SOx and NOx * Soil acidification, buildup of hydrogen cations, which reduces the ...
. The other threat to anemonefish is collection for the marine aquarium trade where anemonefish make up 43% of the global marine ornamental trade, and 25% of the global trade comes from fish bred in captivity, while the majority are captured from the wild, accounting for decreased densities in exploited areas. While bleaching is a significant threat to anemonefish and their host anemones, there is evidence suggesting that collection compounds the localised impact of bleaching. This species was not evaluated in the 2012 release of the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biol ...


In aquaria

The species has successfully been bred in an aquarium. It may move or cover corals with sand. ''A. melanopus'' may become territorial and aggressive once established in a tank. This fish will accept a variety of foods in the aquarium, including flakes, brine shrimp, mysid shrimp, algae, and pellets.


References


External links

* * * * {{Taxonbar, from=Q253437 Amphiprion Fish described in 1852