''Odocoileus lucasi'' (historically ''Navahoceros fricki''), known commonly as the American mountain deer, is an extinct species of North American
deer
Deer or true deer are hoofed ruminant mammals forming the family Cervidae. The two main groups of deer are the Cervinae, including the muntjac, the elk (wapiti), the red deer, and the fallow deer; and the Capreolinae, including the re ...
.
Taxonomy
![Navahoceros skeleton](https://upload.wikimedia.org/wikipedia/commons/8/88/Navahoceros_skeleton.jpg)
Kurten described a species he called ''Navahoceros fricki'' in 1975. However, his analysis has been questioned on technical grounds and new paleontological data. Kurten's analysis was based on averages of length of dissociated bones (samples sizes 9–52), without specifying the sex or age of the source animals, and without providing standard deviations to let the reader know about variability due to sex and age.
[
One comparative element he used was the skeletal measurements of a single mule deer, but he did not provide the data on sex, age or locality. However, from data provided by Klein (1964)][ and McMahon (1975),][ the relative lower leg length of mule deer can vary at least by 22%.
Kurten also stated that ''Navahoceros'' resembled ibex (''Capra ibex'') based on short metapodials which made him label the ''Navahoceros'' as a mountain deer with an Alpine climbing mode of locomotion (like ibex), but without providing data on ibex.][ However, data from Fernandez and Monchot (2007)][ on ibex show that their bone measurements are far from the averages of ''Navahoceros'' presented by Kurten.
Kurten made the explicit correlation that '']Hippocamelus
''Hippocamelus'' is a genus of Cervidae, the deer family. It comprises two extant Andean and two fossil species. The living members are commonly known as the huemul (from the Mapuche language), and the taruca.
Both species have a stocky, thick ...
'' was related to ''Navahoceros'', only differing by having two, instead of three antler tines, and he thus considered ''Hippocamelus'' implicitly to be homologous to chamois and ibex.[ However, ''Hippocamelus'' grow 4 and even 5 tines on each antler, invalidating Kurten's claim, and regarding skeletal proportion, he provided no data on ''Hippocamelus''. Recently, complete appendages were compared between South Andean deer (''Hippocamelus bisulcus''), mountain goats (''Oreamnos americanus''), ibex, Himalayan Tahr (''Hemitragus jemlahicus''), bighorn sheep (''Ovis canadensis''), ''Navahoceros'', chamois (''Rupicapra rupicapra''), mountain sheep (''Ovis ammon''), tule elk (''Cervus canadensis nannodes''), mule deer (''Odocoileus hemionus''), ''Odocoileus lucasi'', red deer (''Cervus elaphus''), and blackbuck (''Antilope cervicapra'').][ Huemul morphology did not overlap with rock climbing species previously considered analogous, but falls within the range of other cervids. In fact, considering the reported variation on leg proportions among several cervids, which can reach 70%, there are ''Rangifer'' and ''Odocoileus virginianus'' populations with shorter legs than the ''H. bisulcus'' sample (by 14%).
Morejohn and Dailey (2004)][ published the analysis of the osteological anatomy and morphology of a practically complete skeleton of a ]Pleistocene
The Pleistocene ( , often referred to as the ''Ice age'') is the geological epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed in ...
adult male ''Odocoileus lucasi'' (Hay 1927) along with other collections labeled as ''O. lucasi''. Moreover, for their 54-page analysis they visited most collections of samples identified as ''Navahoceros'' as well as other species for a comparative analysis (''Cervalces scotti, Alces alces, Rangifer tarandus, Odocoileus hemionus, O. virginianus, Hippocamelus antisensis, H. bisulcus, Mazama americana, Pudu mephistophiles, P. puda, Ozotoceros bezoarticus, Blastocerus dichotomus''), located in 27 different institutions worldwide. They also dissected and analyzed fresh materials of ''Alces, Cervus, Mazama, Odocoileus hemionus, O. virginianus, Ozotoceros, Pudu'', and ''Rangifer''.
Their main conclusions are that ''Navahoceros'' is a '' nomen nudum'' and all revised bones which were labeled as ''Navahoceros'' belong to ''Odocoileus lucasi'' (including exhibition mounts assembled from dissociated bones).[
Although ''Navahoceros'' entered the scene based on a very weak analysis which is impossible to verify,][ the best current evidence based on an extensive comparative study shows that ''Navahoceros'' was an invalid construct and pertains to ''Odocoileus lucasi'' Hay 1927.][ Subsequent publications referring to Kurten's ''Navahoceros'' simply cited his interpretations without questioning its validity. It is recommended that any future discussion, or reference to ''Navahoceros'', be done explicitly in relation to the known information on ''Odocoileus lucasi''.
The interpretations given above are not without controversy. Morejohn and Dailey (2004) were primarily focused on documenting differences between Old World cervids and those of the New World. The fossil skeletal material from Honey Lake was assigned to ''Odocoileus lucasi'' on perceived similarities with the holotype, a first phalanx, and the paratype, an astragalus.][ However, no discussion of the perceived similarities was given and, in the discussion regarding the first phalanx, the only direct comparison mentioned was that of a difference between the Honey Lake first phalanx and that of ''Odocoileus''. It can legitimately be argued insofar as published material goes that it is ''O. lucasi'' that is a ''nomen nudum''.
Other evidence strongly suggests differences between ''Navahoceros'' and ''Odocoileus''. Webb (1992)][ studied a cranium of ''Navahoceros'' from San Josecito Cave and concluded that ''Rangifer'', not ''Odocoileus'', was the sister taxon of ''Navahoceros''. Both ''Blastoceras'' and ''Hippocamelus'' were found to be closer to ''Navahoceros'' than the latter was to other members of the subfamily such as ''Odocoileus''. It is thus far from settled that ''Navahoceros'' is a synonym of ''Odocoileus''.
Assuming that the assignment to ''Odocoileus'' were to be accepted, it is unlikely that the late Pleistocene taxon is conspecific with ''O. lucasi'' (nor did Morejohn and Dailey claim so][). Measurements from the literature and the geologic time span involved would indicate otherwise.
]
Biology
Based on size comparison with modern mule deer the adult, articulated, Honey Lake male specimen would have weighed approximately , significantly larger than the modern species. Hay's original type specimens (toe element and astragalus) were so large that Hay originally placed it in the genus ''Cervus''; the elements are about the same size as those of the Tule elk, the smallest subspecies of ''Cervus elaphus''. The direct anatomical comparison of Hay's two specimens from Idaho and the Honey Lake, California deer by Morejohn confirmed they were conspecific (though not explicitly stated in the text) and both are from a large odocoileine deer.
Range
Kurten in 1975 described ''Navahoceros fricki'' as an extinct member of the family Cervidae and was most common in the North American Rocky Mountains
The Rocky Mountains, also known as the Rockies, are a major mountain range and the largest mountain system in North America. The Rocky Mountains stretch in straight-line distance from the northernmost part of western Canada, to New Mexico ...
during the Pleistocene
The Pleistocene ( , often referred to as the ''Ice age'') is the geological epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed in ...
. It survived to about 11,500 BP from evidence found in Burnet Cave
Burnet Cave (also known as Rocky Arroyo Cave of Wetmore) is an important archaeological and paleontological site located in Eddy County, New Mexico, United States within the Guadalupe Mountains about 26 miles west of Carlsbad.
Physical details
...
in the Guadalupe Mountains
The Guadalupe Mountains ( es, Sierra de Guadalupe) are a mountain range located in West Texas and southeastern New Mexico. The range includes the highest summit in Texas, Guadalupe Peak, , and the "signature peak" of West Texas, El Capitan, both ...
of southern New Mexico
)
, population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano)
, seat = Santa Fe
, LargestCity = Albuquerque
, LargestMetro = Tiguex
, OfficialLang = None
, Languages = English, Spanish ( New Mexican), Navajo, Ke ...
.[
]
References
Further reading
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{{Taxonbar, from=Q7077903
Odocoileus
Prehistoric deer
Pleistocene even-toed ungulates
Pleistocene mammals of North America
Deer Mountain
Fossil taxa described in 1927