The Iberian midwife toad or brown midwife toad (''Alytes cisternasii''), in Portuguese ''sapo-parteiro-ibérico'', is a species of

frog A frog is any member of a diverse and largely Carnivore, carnivorous group of short-bodied, tailless amphibians composing the order (biology), order Anura (ανοὐρά, literally ''without tail'' in Ancient Greek). The oldest fossil "proto-f ...

in the family

Alytidae The Alytidae are a family (biology), family of primitive frogs. Their common name is painted frogs or midwife toads. Most are Endemism, endemic to Europe, but three species occur in northwest Africa, and a species formerly thought to be extinct i ...

(formerly Discoglossidae) found in

Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of ...

and western

Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...

. It is typically found in open habitats such as meadows and open oak forests.

Habitat loss Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...

is one of the threats to its survival.

Description

The Iberian midwife toad grows to a length of about , males being rather smaller than females. The snout is rounded and the eyes large, with vertical slit pupils. Tiny, often orange, warts occur on the upper eyelids. The

parotoid gland The parotoid gland (alternatively, paratoid gland) is an external skin gland on the back, neck, and shoulder of toads and some frogs and salamanders. It can secrete a number of milky alkaloid substances (depending on the species) known collective ...

s are relatively small and the

tympani Timpani (; ) or kettledrums (also informally called timps) are musical instruments in the percussion family. A type of drum categorised as a hemispherical drum, they consist of a membrane called a head stretched over a large bowl traditionally ...

are distinct. Many tubercles are seen on the body and concentrations of glandular warts are under the arms, in the groin area, and on the ankles. The limbs are fairly short. The colour of the upper surface is brownish-grey with dark spots, and the warts are often reddish. The underparts are unspotted and greyish-white.

Distribution and habitat

This toad is native to Portugal and western Spain at altitudes up to above sea level. Its preferred habitat is Mediterranean-type scrub, rough grazing, and light oak woodland.

Biology

Mating takes place in the autumn and the eggs are laid on land. The male then gathers up the egg mass and wraps it round his legs, carrying it around until the developing embryos are ready to hatch. He can carry as many as 180 eggs resulting from four clutches laid by different females. The male deposits the hatching tadpoles in suitable water bodies where they continue their development.

Metamorphosis Metamorphosis is a biological process by which an animal physically develops including birth or hatching, involving a conspicuous and relatively abrupt change in the animal's body structure through cell growth and differentiation. Some inse ...

occurs about four months later when the tadpoles measure about in length.

Status

The main threats it faces are the degradation of suitable terrestrial habitat, pollution, loss of suitable breeding pools, and the introduction of the invasive crayfish ''

Procambarus clarkii ''Procambarus clarkii'', known variously as the red swamp crayfish, Louisiana crawfish or mudbug, is a species of cambarid crayfish native to freshwater bodies of northern Mexico, and southern and southeastern United States, but also introduc ...

'' and non-native fishes that prey on the

tadpole A tadpole is the larval stage in the biological life cycle of an amphibian. Most tadpoles are fully aquatic, though some species of amphibians have tadpoles that are terrestrial. Tadpoles have some fish-like features that may not be found i ...

s. It is also threatened by the infectious fungal disease

chytridiomycosis Chytridiomycosis ( ) is an infectious disease in amphibians, caused by the chytrid fungi ''Batrachochytrium dendrobatidis'' and ''Batrachochytrium salamandrivorans''. Chytridiomycosis has been linked to dramatic population declines or extincti ...

References

External links

* * {{Taxonbar, from=Q927501 Alytes Endemic amphibians of the Iberian Peninsula Amphibians described in 1879 Taxa named by Eduardo Boscá Taxonomy articles created by Polbot