An alvar is a biological environment based on a limestoneplain with thin or no soil and, as a result, sparse grassland vegetation. Often flooded in the spring, and affected by drought in midsummer, alvars support a distinctive group of prairie-like plants. Most alvars occur either in northern Europe or around the Great Lakes in North America. This stressed habitat supports a community of rare plants and animals, including species more commonly found on prairiegrasslands.
Lichen
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.moss
Mosses are small, non-vascular flowerless plants in the taxonomic division Bryophyta (, ) '' sensu stricto''. Bryophyta (''sensu lato'', Schimp. 1879) may also refer to the parent group bryophytes, which comprise liverworts, mosses, and hor ...
es are common species. Trees and bushes are absent or severely stunted.
The primary cause of alvars is the shallow exposed bedrock. Flooding and drought, as noted, add to the stress of the site and prevent many species from growing. Disturbance may also play a role. In Europe, grazing is frequent, while in North America, there is some evidence that fire may also prevent encroachment by forest. The habitat also has strong competition gradients, with better competitors occupying the deeper soil and excluding other species to less productive locations. Crevices in the limestone provide a distinctive habitat which is somewhat protected from grazing, and which may provide habitat for unusual ferns such as '' Pellaea atropurpurea''. Bare rock flats provide areas with extremely low competition that serve as refugia for weak competitors such as the sandwort ''
Minuartia michauxii
''Minuartia'' is a genus of flowering plants commonly known as sandworts in the family Caryophyllaceae.
Minuartias are small annual or perennial plants which grow in otherwise inhospitable conditions such as on rocky ledges and in stony soil. Th ...
'' and '' Micranthes virginiensis''. In a representative set of four Ontario alvars, seven habitat types were described. From deep to shallow soil these were: tall grassy meadows, tall forb-rich meadows, low grassy meadows, low forb-rich meadows, dry grassland, rock margin grassland and bare rock flats.
Alvars comprise a small percentage of the Earth's ecosystems by land extent. Although some 120 exist in the Great Lakes region, in total there are only about left across the entire Great Lakes basin, and many of these have been degraded by agriculture and other human uses. More than half of all remaining alvars occur in Ontario. There are smaller areas in New York, Michigan, Ohio, Wisconsin and Quebec.
In
North America
North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
butterflies
Butterflies are insects in the macrolepidopteran clade Rhopalocera from the Order (biology), order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The ...
and snails. The use of the word "alvar" to refer to this type of environment originated in
Sweden
Sweden, formally the Kingdom of Sweden,The United Nations Group of Experts on Geographical Names states that the country's formal name is the Kingdom of SwedenUNGEGN World Geographical Names, Sweden./ref> is a Nordic country located on ...
. The largest alvar in Europe is located on the Swedish island of Öland. Here the thin soil mantle is only 0.5 to 2.0 centimeters thick in most places and in many extents consists of exposed limestone slabs. The landscape there has been designated a UNESCOWorld Heritage Site. There are other more local names for similar landforms, such as a pavement barren, although this term is also used for similar
landform
A landform is a natural or anthropogenic land feature on the solid surface of the Earth or other planetary body. Landforms together make up a given terrain, and their arrangement in the landscape is known as topography. Landforms include hills, ...
s based on sandstone. In the United Kingdom the exposed landform is called a
limestone pavement
A limestone pavement is a natural karst landform consisting of a flat, incised surface of exposed limestone that resembles an artificial pavement. The term is mainly used in the UK and Ireland, where many of these landforms have developed dist ...
Sweden
Sweden, formally the Kingdom of Sweden,The United Nations Group of Experts on Geographical Names states that the country's formal name is the Kingdom of SwedenUNGEGN World Geographical Names, Sweden./ref> is a Nordic country located on ...
Gotland
Gotland (, ; ''Gutland'' in Gutnish), also historically spelled Gottland or Gothland (), is Sweden's largest island. It is also a province, county, municipality, and diocese. The province includes the islands of Fårö and Gotska Sandön to the ...
** Västergötland – several locations on limestone mountain
Kinnekulle
Kinnekulle is a flat-topped mountain in the county of Västergötland, Sweden, on the eastern shore of lake Vänern. Its highest point is above sea level. The mountain is long and wide at the top.
Geology
Despite its enormous size, Kinnekul ...
, smaller fragments on
Falbygden
Falbygden is a geographical area, centered at the town of Falköping in Västergötland,
in southwestern Sweden, and covered mostly by farmland. Most of the area belongs to Falköping Municipality and the west part of Tidaholm Municipality. In med ...
, e.g. in Dala and Högstena parishes
* Estonia
** Alvars are distributed along the whole northern coast of Estonia from approx. the town of
Paldiski
Paldiski is a town and Baltic Sea port situated on the Pakri Peninsula of northwestern Estonia. Since 2017, it's the administrative centre of Lääne-Harju Parish of Harju County. Previously a village of Estonia-Swedes known by the historical ...
to Sillamäe, wherever limestone comes to the surface near the seashore (see Baltic Klint), as well as on the islands of the West Estonian archipelago. Estonia used to be home to approximately one third of the world's alvars; however, the total area of alvars has decreased from 43,000 hectares in the 1930s to 12,000 hectares in 2000, and approximately 9,000 hectares in 2010. Estonian alvars are home to 267 species of vascular plants, approximately one fifth of which are protected. There are also 142 species of bryophytes and 263 species of
lichen
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship. The Estonian government has committed itself to protect at least 9,800 hectares of the country's alvars as part of the
Natura 2000
Natura 2000 is a network of nature protection areas in the territory of the European Union. It is made up of Special Areas of Conservation and Special Protection Areas designated under the Habitats Directive and the Birds Directive, respectively ...
network. The
Loopealse
Loopealse ( Estonian for ''"Alvar"'', literally ''"Above Alvar"'') is a subdistrict ( et, asum) in the district of Lasnamäe, Tallinn, the capital of Estonia. It has a population of 2,388 ().
Gallery
File:Tallinn Loopealse view.jpg,
File:EE- ...
Vardi Nature Reserve
Vardi Nature Reserve is a nature reserve in Rapla County in central Estonia.
Vardi nature reserve encompasses one of the few remaining alvar forests in Estonia, an unusual landscape type today as cut-down forests normally never recover. The soi ...
in Rapla County is an Estonian nature reserve especially designated to protect one of the more representative alvar areas of Estonia.
* England
** Cumbria and North Yorkshire – under protection in the UK Biodiversity Action Plan
* Ireland
** The Burren, a large alvar in northwest County Clare
Sarve loopealne Hiiumaal.jpg, Sarve alvar in Hiiumaa island
Väike-Pakri loopealsel.jpg, Alvar in
island
Österplana hed vegetation.jpg, Sparse vegetation,
Kinnekulle
Kinnekulle is a flat-topped mountain in the county of Västergötland, Sweden, on the eastern shore of lake Vänern. Its highest point is above sea level. The mountain is long and wide at the top.
Geology
Despite its enormous size, Kinnekul ...
,
Sweden
Sweden, formally the Kingdom of Sweden,The United Nations Group of Experts on Geographical Names states that the country's formal name is the Kingdom of SwedenUNGEGN World Geographical Names, Sweden./ref> is a Nordic country located on ...
Limestone pavement above Malham Cove.jpg, Limestone pavement at Malham Cove, UK
Kelley's Island
Kelleys Island is both a village in Erie County, Ohio, and the island which it fully occupies in Lake Erie. The British originally called it Sandusky Island. Later the United States took it over and officially designated it as Island Number 6 ...
, Ohio – North Shore Alvar State Nature Preserve
** Marblehead, Ohio – mostly destroyed by limestone quarrying
** Pelee Island, Ontario – Stone Road Alvar Nature Reserve
*
Lake Huron
Lake Huron ( ) is one of the five Great Lakes of North America. Hydrology, Hydrologically, it comprises the easterly portion of Lake Michigan–Huron, having the same surface elevation as Lake Michigan, to which it is connected by the , Strait ...
**Maxton Plains Proposed Natural Area, Drummond Island, Michigan
**Belanger Bay Alvar, Manitoulin Island, Ontario
**Quarry Bay Nature Reserve, Manitoulin Island, Ontario
**Bruce Alvar Nature Reserve, Bruce Peninsula, Ontario
**Baptise Harbour Nature Reserve, Bruce Peninsula, Ontario
**
Misery Bay
Misery Bay is a freshwater bay on Manitoulin Island in Ontario. It is noted for its alvars and large dolomite pavements ground flat by glaciers. It also contains grykes and ecologically significant wetlands.
History
Misery Bay was covered b ...
Provincial Park, Manitoulin Island, Ontario
*
Lake Michigan
Lake Michigan is one of the five Great Lakes of North America. It is the second-largest of the Great Lakes by volume () and the third-largest by surface area (), after Lake Superior and Lake Huron. To the east, its basin is conjoined with that o ...
**Red Banks Alvar,
Red Banks, Brown County, Wisconsin
Red Banks is an unincorporated community located in the town of Scott, Brown County, Wisconsin, United States.
History
In 1634, Jean Nicolet
Jean Nicolet (Nicollet), Sieur de Belleborne (October 1642) was a French '' coureur des bois'' note ...
* Lake Ontario
**Carden Plain Alvar, City of Kawartha Lakes, Ontario, including
Carden Alvar Provincial Park
Carden Alvar Provincial Park is a provincial park located in the Kawartha Lakes in Central Ontario, Canada. Alvars are globally rare ecosystems found exclusively in Northern Europe and the Great Lakes region of North America. The park is classif ...
**Chaumont Barrens Preserve, New York
**Three Mile Creek Barrens, New York
**
Burnt Lands Alvar
The Burnt Lands is an alvar between Almonte and Ottawa near Upper Huntley, Ontario, Canada. It probably obtained its name from one of the forest fires that swept the area during early European settlement.
It is possible that fires assist in crea ...
Quyon
Quyon is a village that is part of Pontiac, Quebec, in the Les Collines-de-l'Outaouais Regional County Municipality (MRC des Collines).
History
Already the site of the Sainte-Marie Mission, the village was founded in 1848 by John Egan, a lumb ...
Interlake
Interlake was a provincial electoral division in the Canadian province of Manitoba. It was created by redistribution in 1979, and has formally existed since the 1981 provincial election. Previously, much of the Interlake region was include ...