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The alula , or bastard wing, (plural ''alulae'') is a small projection on the
anterior Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position prov ...
edge of the
wing A wing is a type of fin that produces lift while moving through air or some other fluid. Accordingly, wings have streamlined cross-sections that are subject to aerodynamic forces and act as airfoils. A wing's aerodynamic efficiency is expres ...
of modern
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
s and a few
non-avian dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
s. The word is
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
and means "winglet"; it is the
diminutive A diminutive is a root word that has been modified to convey a slighter degree of its root meaning, either to convey the smallness of the object or quality named, or to convey a sense of intimacy or endearment. A (abbreviated ) is a word-formati ...
of ''ala'', meaning "wing". The alula is the freely moving first digit, a bird's "thumb", and typically bears three to five small
flight feather Flight feathers (''Pennae volatus'') are the long, stiff, asymmetrically shaped, but symmetrically paired pennaceous feathers on the wings or tail of a bird; those on the wings are called remiges (), singular remex (), while those on the tail ...
s, with the exact number depending on the
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
. There also are minor
covert feather A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which, as the name implies, cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are sm ...
s overlying the flight feathers. Like the larger
flight feather Flight feathers (''Pennae volatus'') are the long, stiff, asymmetrically shaped, but symmetrically paired pennaceous feathers on the wings or tail of a bird; those on the wings are called remiges (), singular remex (), while those on the tail ...
s found on the wing's trailing edge, these alula feathers are asymmetrical, with the shaft running closer to anterior edge.


Function

In most situations, the alula is held flush against the wing; however, it can be manipulated. When flying at slow speeds or landing, the bird moves its alula slightly upwards and forward, which creates a small slot on the wing's leading edge. This functions in the same way as the
slats Super Low Altitude Test Satellite (SLATS) or Tsubame was a JAXA satellite intended to demonstrate operations in very low Earth orbit (VLEO, below 200 km), using ion engines to counteract aerodynamic drag from the Earth's atmosphere which i ...
on the wing of an aircraft, allowing the wing to achieve a higher than normal
angle of attack In fluid dynamics, angle of attack (AOA, α, or \alpha) is the angle between a reference line on a body (often the chord line of an airfoil) and the vector representing the relative motion between the body and the fluid through which it is m ...
– and thus
lift Lift or LIFT may refer to: Physical devices * Elevator, or lift, a device used for raising and lowering people or goods ** Paternoster lift, a type of lift using a continuous chain of cars which do not stop ** Patient lift, or Hoyer lift, mobile ...
– without resulting in a stall. The tip of the alula forms a tiny vortex, acting similar to a
vortex generator A vortex generator (VG) is an aerodynamic device, consisting of a small vane usually attached to a lifting surface (or airfoil, such as an aircraft wing) or a rotor blade of a wind turbine.peregrine falcon The peregrine falcon (''Falco peregrinus''), also known as the peregrine, and historically as the duck hawk in North America, is a Cosmopolitan distribution, cosmopolitan bird of prey (Bird of prey, raptor) in the family (biology), family Falco ...
s.


In ancient birds

The presence of an alula has been confirmed in several now-
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
ancient relatives of modern birds, including ''
Eoalulavis hoyasi ''Eoalulavis'' (from the Ancient Greek: ''Éōs'', "dawn"; ''alula'', "bastard wing"; ''avis'', "bird") is a monotypic genus of enantiornithean bird that lived during the Barremian, in the Lower Cretaceous around 125 million years ago. The only ...
'' (an
enantiornithine The Enantiornithes, also known as enantiornithines or enantiornitheans in literature, are a group of extinct avialans ("birds" in the broad sense), the most abundant and diverse group known from the Mesozoic era. Almost all retained teeth and cla ...
from the mid-
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...
, 115 mya) and the earlier ''
Protopteryx fengningensis ''Protopteryx'' is an extinct bird and the most basal enantiornithean, from the Cretaceous period.Zhou, Zhonghe. (2002). "A new and primitive enantiornithine bird from the Early Cretaceous of China." ''Journal of Vertebrate Paleontology'', 22(1): ...
''. Since these
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
are not closely related to modern birds, either the alula
evolved Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation t ...
twice, or it did so more than 130 million years ago.


See also

* Alular digit homology *
Bird flight Bird flight is the primary mode of locomotion used by most bird species in which birds take off and fly. Flight assists birds with feeding, breeding, avoiding predators, and migrating. Bird flight is one of the most complex forms of locomotion ...
*
Leading edge slot A leading-edge slot is a fixed aerodynamic feature of the wing of some aircraft to reduce the stall speed and promote good low-speed handling qualities. A leading-edge slot is a spanwise gap in each wing, allowing air to flow from below the wing ...
, the man-made equivalent on fixed-wing aircraft


Citations


References

* * * {{Birds Feathers Bird anatomy