Alternate-leaved Dogwood
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''Cornus alternifolia'' is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
in the dogwood
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Cornaceae The Cornaceae are a cosmopolitan family of flowering plants in the order Cornales. The family contains approximately 85 species in two genera, ''Alangium'' and ''Cornus''. They are mostly trees and shrubs, which may be deciduous or evergreen, alt ...
,
native Native may refer to: People * Jus soli, citizenship by right of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Native Americans (disambiguation) In arts and entert ...
to eastern North America, from Newfoundland west to southern Manitoba and Minnesota, and south to northern Florida and Mississippi. It is rare in the southern United States. It is commonly known as green osier, alternate-leaved dogwood, and pagoda dogwood.


Description

It is a small
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, aft ...
shrub or tree growing to (rarely ) tall, with a trunk up to in diameter. The branches develop characteristic horizontal layers separated by gaps, with a flat-topped crown. Its
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
are elliptic to ovate and grow to long and broad, arranged alternately on the stems, not in opposite pairs typical of the majority of ''Cornus'' species. The leaves are most often arranged in crowded clusters around the ends of the twigs and appear almost whorled. The upper sides of the leaves are smooth and green, while the undersides are hairy and a bluish color. The
bark Bark may refer to: * Bark (botany), an outer layer of a woody plant such as a tree or stick * Bark (sound), a vocalization of some animals (which is commonly the dog) Places * Bark, Germany * Bark, Warmian-Masurian Voivodeship, Poland Arts, ...
is colored gray to brown, becoming ridged as it ages. Small cream colored
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...
s are produced, with four small petals. The flowers are grouped into cymes, with the
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...
s across. It bears
berries A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit, although many pips or seeds may be present. Common examples are strawberries, raspb ...
with a blackish blue color. * Bark: Dark reddish brown, with shallow ridges. Branchlets at first pale reddish green, later dark green. * Wood: Reddish brown, sapwood pale; heavy, hard, close-grained. Sp. gr., 0.6696; weight . * Winter buds: Light chestnut brown, acute. Inner scales enlarge with the growing shoot and become half an inch long before they fall. * Leaves: Alternate, rarely opposite, often clustered at the ends of the branch, simple, three to five inches long, two to three wide, oval or ovate, wedge-shaped or rounded at base; margin is wavy toothed, slightly reflexed, apex acuminate. They come out of the bud involute, reddish green above, coated with silvery white tomentum beneath, when full grown are bright green above, pale, downy, almost white beneath. Feather-veined, midrib broad, yellowish, prominent beneath, with about six pairs of primary veins. In autumn they turn yellow, or yellow and scarlet. Petioles slender, grooved, hairy, with clasping bases. * Flowers: April, May. Perfect, cream color, borne in many-flowered, broad, open cymes, at the end of short lateral branches. * Calyx: The cup-shaped flowers have four petals that are valvate in bud, unwrapping when in bloom with cream colored, oblong shaped petals with rounded ends. The petals are inserted on disk and the stamens are inserted too and arranged alternately to the petals, being four in number also. The stamens are exserted with filaments long and slender. Anthers oblong, introrse, versatile, two-celled; cells opening longitudinally. * Pistil: Ovary inferior, two-celled; style columnar; stigma capitate. * Fruit: Drupe, globular, blue-black, across, tipped with remnant of style which rises from a slight depression; nut obovoid, many-grooved. October.


Habitat

''C. alternifolia'' is found under open deciduous trees, as well as along the margins of forests and swamps. These trees prefer moist, well drained soil. Seedlings are shade-tolerant and it is often found as an understory tree in mature forests, such as those dominated by ''
Acer saccharum ''Acer saccharum'', the sugar maple, is a species of flowering plant in the soapberry and lychee family Sapindaceae. It is native to the hardwood forests of eastern Canada and eastern United States. Sugar maple is best known for being the prima ...
'' (sugar maple) or ''
Populus ''Populus'' is a genus of 25–30 species of deciduous flowering plants in the family Salicaceae, native to most of the Northern Hemisphere. English names variously applied to different species include poplar (), aspen, and cottonwood. The we ...
'' (aspen). It is also common in younger forests.


Ecology

The fruits provide food for at least eleven species of birds and the
black bear Black bear or Blackbear may refer to: Animals * American black bear (''Ursus americanus''), a North American bear species * Asian black bear (''Ursus thibetanus''), an Asian bear species Music * Black Bear (band), a Canadian First Nations group ...
. The leaves and bark are eaten by
white-tailed deer The white-tailed deer (''Odocoileus virginianus''), also known as the whitetail or Virginia deer, is a medium-sized deer native to North America, Central America, and South America as far south as Peru and Bolivia. It has also been introduced t ...
,
beaver Beavers are large, semiaquatic rodents in the genus ''Castor'' native to the temperate Northern Hemisphere. There are two extant species: the North American beaver (''Castor canadensis'') and the Eurasian beaver (''C. fiber''). Beavers ar ...
, and
cottontail rabbit Cottontail rabbits are the leporid species in the genus ''Sylvilagus'', found in the Americas. Most ''Sylvilagus'' species have stub tails with white undersides that show when they retreat, giving them their characteristic name. However, this ...
. It additionally has been marked as a pollinator plant, supporting and attracting bees, flies, and butterflies. It is a host plant for butterflies, providing food during their larval stage.


Use

The tree is regarded as attractive because of its wide-spreading shelving branches and flat-topped head, and is often used in ornamental plantings. The flower clusters have no great white
involucre In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves. They may be smaller, larger, or of ...
as have those of the
flowering dogwood ''Cornus florida'', the flowering dogwood, is a species of flowering tree in the family Cornaceae native to eastern North America and northern Mexico. An endemic population once spanned from southernmost coastal Maine south to northern Florida ...
, and the fruit is dark purple instead of red. The
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture, ...
'Argentea' (silver pagoda dogwood) has gained the
Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr (Nort ...
's
Award of Garden Merit The Award of Garden Merit (AGM) is a long-established annual award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. History The Award of Garden Merit ...
(confirmed 2017). ''C. alternifolia'' is susceptible to golden canker ('' Cryptodiaporthe corni''), particularly when drought-stressed or heat-stressed. Proper siting of the plant in partial to full shade, along with adequate mulch and water, will reduce the incidence of this
pathogen In biology, a pathogen ( el, πάθος, "suffering", "passion" and , "producer of") in the oldest and broadest sense, is any organism or agent that can produce disease. A pathogen may also be referred to as an infectious agent, or simply a germ ...
. ''Cornus alternifolia'' has been used in the
traditional Chinese medicine Traditional Chinese medicine (TCM) is an alternative medical practice drawn from traditional medicine in China. It has been described as "fraught with pseudoscience", with the majority of its treatments having no logical mechanism of action ...
as tonic,
analgesic An analgesic drug, also called simply an analgesic (American English), analgaesic (British English), pain reliever, or painkiller, is any member of the group of drugs used to achieve relief from pain (that is, analgesia or pain management). It ...
, and
diuretic A diuretic () is any substance that promotes diuresis, the increased production of urine. This includes forced diuresis. A diuretic tablet is sometimes colloquially called a water tablet. There are several categories of diuretics. All diuretics in ...
.


Gallery

Jk-cornus alternifoliaflower.jpg, Pagodaseedlings.jpg, Cornus alternifolia 005.jpg,


References


Further reading

#''Trees'', by Coombes, Allen J., Eyewitness Handbooks


External links


Natural Resources Canada: ''Cornus alternifolia''


{{Taxonbar, from=Q639615 alternifolia Medicinal plants of North America Flora of Eastern Canada Flora of the North-Central United States Trees of the Northeastern United States Trees of the Southeastern United States Trees of Manitoba