Alfaroa Columbiana
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''Alfaroa'' is a genus of evergreen trees in the
Juglandaceae The Juglandaceae are a plant family known as the walnut family. They are trees, or sometimes shrubs, in the order Fagales. Members of this family are native to the Americas, Eurasia, and Southeast Asia. The nine or ten genera in the family have ...
family of the
Fagales The Fagales are an order of flowering plants, including some of the best-known trees. The order name is derived from genus ''Fagus'', beeches. They belong among the rosid group of dicotyledons. The families and genera currently included are a ...
, growing in montane and submontane tropical rain forests in Central America. The wood is characterized by solid pith, pink heartwood, and vessels with scalariform perforations, as well as simple perforations.


Description

The (usually
pinnate Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and in ...
ly compound) leaves are evergreen and lack
stipule In botany, a stipule is an outgrowth typically borne on both sides (sometimes on just one side) of the base of a leafstalk (the petiole). Stipules are considered part of the anatomy of the leaf of a typical flowering plant, although in many speci ...
s. They are alternate, rarely opposite. The plants are monoecious, the male flowers being in lateral panicles (several pairs of catkins on an inflorescence) and the female flowers born terminally either in a single spike or in a hermaphroditic panicle including several paired male catkins. Each flower has a wide bract, two bracteoles, and four sepals. The flowers are sessile. The male flowers have a round or oblong receptacle and six to ten stamens. The pollen grains are approximately 24 micrometers in diameter and are slightly triangular in polar view. The small fruits are nuts, one-chambered at the apex and eight-chambered (sometimes four-chambered) at the base. Germination is
hypogeal Hypogeal, hypogean, hypogeic and hypogeous (; ) are biological terms describing an organism's activity below the soil surface. In botany, a seed is described as showing hypogeal germination when the cotyledons of the germinating seed remain n ...
.


Species

''Alfaroa'' includes the following species (This list may be incomplete): * '' A. columbiana'', G. Lozano-C., J. Hernandez-C., & S. Espinal-T. * '' A. costaricensis'',
Standl. Paul Carpenter Standley (March 21, 1884 – June 2, 1963) was an American botanist known for his work on neotropical plants. __TOC__ Standley was born on March 21, 1884 in Avalon, Missouri. He attended Drury College in Springfield, Missouri an ...
—Campano Chile, Chiciscua, Gaulin, Gavilancillo * '' A. guanacastensis'', D. E. Stone * '' A. guatemalensis'', (
Standl. Paul Carpenter Standley (March 21, 1884 – June 2, 1963) was an American botanist known for his work on neotropical plants. __TOC__ Standley was born on March 21, 1884 in Avalon, Missouri. He attended Drury College in Springfield, Missouri an ...
) L. O. Williams & A. Molina 1970 * '' A. hondurensis'', L.O. Williams 1959 * '' A. manningii'', Jorge Leon * '' A. mexicana'', D. E. Stone * '' A. roxburghiana'' (
Wall. Nathaniel Wolff Wallich FRS FRSE (28 January 1786 – 28 April 1854) was a surgeon and botanist of Danish origin who worked in India, initially in the Danish settlement near Calcutta and later for the Danish East India Company and the British ...
) I. A. Iljinsk. * '' A. williamsii'', A. Modina R. ''A. guatemalensis'' and ''A. roxburghiana'' were originally described as belonging to the genus ''
Engelhardia ''Engelhardia'' is a genus of seven species of trees in the family Juglandaceae, native to southeast Asia from northern India east to Taiwan, Indonesia and the Philippines. The genus name is commonly misspelled "''Engelhardtia''", a "correctio ...
''. The monotypic genus ''Alfaropsis'' I. A. Iljinsk has also been erected for ''A. roxburghiana''.


References

Elliott, L. L., Mindell, R. A., & Stockey, R. A., "''Beardia vancouverensis'' Gen. et Sp. Nov. (Juglandaceae): Permineralized Fruits from the Eocene of British Columbia", ''American Journal of Botany'' 93(4): 557–565, 2006 Manchester, Steven R., "Fossil Wood of the Engelhardieae (Junglandaceae) from the Eocene of North America: ''Engelhardioxylon'' Gen. Nov.", ''Bot. Gasz'' 144(1)157-163, 1983. J. A. Vozzo (ed.) ''Tropical Tree Seed Manual'' "Part II—Species Descriptions", USDA Forest Service Engelhardioideae Fagales genera Neotropical realm flora Taxonomy articles created by Polbot {{Fagales-stub