Alangium Brachyanthum
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''Alangium'' is a small genus of flowering plants. The genus is included either in a broad view of the dogwood family Cornaceae, or as the sole member of its own family Alangiaceae.Qiu-Yun (Jenny) Xiang, David T. Thomas, and Qiao Ping Xiang. 2011. "Resolving and dating the phylogeny of Cornales - Effects of taxon sampling, data partitions, and fossil calibrations". ''Molecular Phylogenetics and Evolution'' 59(1):123-138. ''Alangium'' has about 40 species, but some of the species boundaries are not entirely clear.Chun-Miao Feng, Steven R. Manchester, and Qiu-Yun (Jenny) Xiang. 2009. "Phylogeny and biogeography of Alangiaceae (Cornales) inferred from DNA sequences, morphology, and fossils". ''Molecular Phylogenetics and Evolution'' 51(2):201-214. The type species for ''Alangium'' is ''Alangium decapetalum'', which is now treated as a
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
of ''Alangium salviifolium''.''Alangium'' In: Index Nominum Genericorum. In: Regnum Vegetabile (see ''External links'' below). All of the species are
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
s or small trees, except the
liana A liana is a long- stemmed, woody vine that is rooted in the soil at ground level and uses trees, as well as other means of vertical support, to climb up to the canopy in search of direct sunlight. The word ''liana'' does not refer to a ta ...
''Alangium kwangsiense''. ''A. chinense, A. platanifolium'', and ''A. salviifolium'' are known in cultivation. Anthony Huxley, Mark Griffiths, and Margot Levy (1992). ''The New Royal Horticultural Society Dictionary of Gardening''. The Macmillan Press,Limited: London. The Stockton Press: New York. (set).


Range

The genus consists of small trees,
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
s and
liana A liana is a long- stemmed, woody vine that is rooted in the soil at ground level and uses trees, as well as other means of vertical support, to climb up to the canopy in search of direct sunlight. The word ''liana'' does not refer to a ta ...
s, and is native to western Africa, Madagascar, southern and eastern Asia (
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
, Malaysia, Indonesia, and the Philippines), tropical
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
, the western Pacific Ocean islands, and
New Caledonia ) , anthem = "" , image_map = New Caledonia on the globe (small islands magnified) (Polynesia centered).svg , map_alt = Location of New Caledonia , map_caption = Location of New Caledonia , mapsize = 290px , subdivision_type = Sovereign st ...
. Most of the species are
native Native may refer to: People * Jus soli, citizenship by right of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Native Americans (disambiguation) In arts and entert ...
to tropical and
subtropical The subtropical zones or subtropics are geographical zone, geographical and Köppen climate classification, climate zones to the Northern Hemisphere, north and Southern Hemisphere, south of the tropics. Geographically part of the Geographical z ...
regions of east and southeast Asia. Five of the species extend well outside of this area. ''Alangium platanifolium'' extends from east Asia into Russia. ''Alangium chinense'' ( sensu lato) extends from southeast Asia to Africa. ''Alangium salviifolium'' is the most widespread species, ranging from Africa to
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
,
Fiji Fiji ( , ,; fj, Viti, ; Fiji Hindi: फ़िजी, ''Fijī''), officially the Republic of Fiji, is an island country in Melanesia, part of Oceania in the South Pacific Ocean. It lies about north-northeast of New Zealand. Fiji consists ...
, and
New Caledonia ) , anthem = "" , image_map = New Caledonia on the globe (small islands magnified) (Polynesia centered).svg , map_alt = Location of New Caledonia , map_caption = Location of New Caledonia , mapsize = 290px , subdivision_type = Sovereign st ...
. ''Alangium villosum'' occurs from southeast Asia to Australia and the western
Pacific Islands Collectively called the Pacific Islands, the islands in the Pacific Ocean are further categorized into three major island groups: Melanesia, Micronesia, and Polynesia. Depending on the context, the term ''Pacific Islands'' may refer to one of se ...
. ''Alangium grisolleoides'' is endemic to Madagascar and gives the genus a
disjunct distribution In biology, a taxon with a disjunct distribution is one that has two or more groups that are related but considerably separated from each other geographically. The causes are varied and might demonstrate either the expansion or contraction of a s ...
. ''Alangium'' species are used as food plants by the larvae of some
Lepidoptera Lepidoptera ( ) is an order (biology), order of insects that includes butterfly, butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 Family (biology), families and 46 Taxonomic r ...
species in the Geometroidea-Drepanoidea assemblage including engrailed ( Geometridae) and the subfamily
Cyclidiinae Cyclidiinae is a small subfamily of the Drepanidae moths. They occur in Southeast Asia. Their caterpillars feed on ''Alangium ''Alangium'' is a small genus of flowering plants. The genus is included either in a broad view of the dogwood f ...
( Drepanidae).


Etymology

The
name A name is a term used for identification by an external observer. They can identify a class or category of things, or a single thing, either uniquely, or within a given context. The entity identified by a name is called its referent. A personal ...
''Alangium'' is a Latinization,
derived Derive may refer to: * Derive (computer algebra system), a commercial system made by Texas Instruments * ''Dérive'' (magazine), an Austrian science magazine on urbanism *Dérive, a psychogeographical concept See also * *Derivation (disambiguatio ...
from the Malayalam
name A name is a term used for identification by an external observer. They can identify a class or category of things, or a single thing, either uniquely, or within a given context. The entity identified by a name is called its referent. A personal ...
''alangi'', which, in Kerala, refers to ''Alangium salviifolium''.Umberto Quattrocchi. 2000. ''CRC World Dictionary of Plant Names'' volume I. CRC Press: Boca Raton; New York; Washington,DC;, USA. London, UK. (vol. I). It was named in 1783 by Jean-Baptiste Lamarck in his '' Encyclopédie Méthodique''.''Alangium'' At: International Plant Names Index. (See ''External links'' below).Jean-Baptiste Lamarck. 1783. ''Encyclopédie Méthodique: botanique''. 1(1):174


Paleontological record

The wood, fruit, and
pollen Pollen is a powdery substance produced by seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametophyt ...
of ''Alangium'' are distinctive. Fossils of ''Alangium'' have been recognized from the early Eocene of England and the middle Eocene of western
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
. In former times, ''Alangium'' was far more widespread than it is today.


Species

''
The Plant List The Plant List was a list of botanical names of species of plants created by the Royal Botanic Gardens, Kew and the Missouri Botanical Garden and launched in 2010. It was intended to be a comprehensive record of all known names of plant species ...
'' recognises 42 accepted species (including infraspecific names): * ''
Alangium alpinum ''Alangium'' is a small genus of flowering plants. The genus is included either in a sensu lato, broad view of the dogwood family Cornaceae, or as the monotypic taxon, sole member of its own Family (biology), family Alangiaceae.Qiu-Yun (Jenny) ...
'' * ''
Alangium barbatum ''Alangium'' is a small genus of flowering plants. The genus is included either in a broad view of the dogwood family Cornaceae, or as the sole member of its own family Alangiaceae.Qiu-Yun (Jenny) Xiang, David T. Thomas, and Qiao Ping Xiang. ...
'' ** subsp. ''barbatum'' ** subsp. ''decipiens'' ** subsp. ''faberi'' * '' Alangium brachyanthum'' * ''
Alangium chinense ''Alangium chinense'' is a species of flowering plant in the Cornaceae family. It has the Chinese name (). Traditional uses It is one of the Chinese herbology#50 fundamental herbs, 50 fundamental herbs used in traditional Chinese medicine. In Hu ...
'' * ''
Alangium chungii ''Alangium'' is a small genus of flowering plants. The genus is included either in a sensu lato, broad view of the dogwood family Cornaceae, or as the monotypic taxon, sole member of its own Family (biology), family Alangiaceae.Qiu-Yun (Jenny) ...
'' * ''
Alangium circulare ''Alangium circulare'' is a tree in the dogwood family Cornaceae. The specific epithet ' is from the Latin meaning "circular", referring to shape of the leaves. Description ''Alangium circulare'' grows as a tree up to tall. The smooth bark is g ...
'' * ''
Alangium grisolleoides ''Alangium'' is a small genus of flowering plants. The genus is included either in a broad view of the dogwood family Cornaceae, or as the sole member of its own family Alangiaceae.Qiu-Yun (Jenny) Xiang, David T. Thomas, and Qiao Ping Xiang. ...
'' * ''
Alangium havilandii ''Alangium havilandii'' is a tree in the dogwood family Cornaceae. It is named for the British surgeon and naturalist George Darby Haviland. Description ''Alangium havilandii'' grows as a tree up to tall with a trunk diameter of up to . The smo ...
'' * ''
Alangium javanicum ''Alangium javanicum'' is a species of plant in the Cornaceae family. It is found in Brunei, Indonesia, Malaysia, Singapore Singapore (), officially the Republic of Singapore, is a sovereign island country and city-state in maritime Sou ...
'' ** var. ''ebenaceum'' ** var. ''papuanum'' * ''
Alangium kurzii ''Alangium kurzii'' is a tree in the family Cornaceae. It is named for the German naturalist Wilhelm Sulpiz Kurz. Description ''Alangium kurzii'' grows up to tall with a trunk diameter of up to . The smooth bark is dark grey. The flowers are pa ...
'' * ''
Alangium longiflorum ''Alangium longiflorum'' is a tree in the dogwood family Cornaceae. The specific name (botany), specific epithet ' is from the Latin meaning "long flowers". Description ''Alangium longiflorum'' grows as a tree up to tall with a trunk diameter o ...
'' * ''
Alangium maliliense ''Alangium'' is a small genus of flowering plants. The genus is included either in a sensu lato, broad view of the dogwood family Cornaceae, or as the monotypic taxon, sole member of its own Family (biology), family Alangiaceae.Qiu-Yun (Jenny) ...
'' * ''
Alangium nobile ''Alangium nobile'' is a tree in the dogwood family Cornaceae. The specific name (botany), specific epithet ' is from the Latin meaning "noble" or "distinguished", likely referring to the growth Glossary of botanical terms#habit, habit. Descript ...
'' * ''
Alangium platanifolium ''Alangium'' is a small genus of flowering plants. The genus is included either in a broad view of the dogwood family Cornaceae, or as the sole member of its own family Alangiaceae.Qiu-Yun (Jenny) Xiang, David T. Thomas, and Qiao Ping Xiang. ...
'' * ''
Alangium premnifolium ''Alangium'' is a small genus of flowering plants. The genus is included either in a sensu lato, broad view of the dogwood family Cornaceae, or as the monotypic taxon, sole member of its own Family (biology), family Alangiaceae.Qiu-Yun (Jenny) ...
'' * ''
Alangium qingchuanense ''Alangium'' is a small genus of flowering plants. The genus is included either in a broad view of the dogwood family Cornaceae, or as the sole member of its own family Alangiaceae.Qiu-Yun (Jenny) Xiang, David T. Thomas, and Qiao Ping Xiang. ...
'' * ''
Alangium ridleyi ''Alangium'' is a small genus of flowering plants. The genus is included either in a broad view of the dogwood family Cornaceae, or as the sole member of its own family Alangiaceae.Qiu-Yun (Jenny) Xiang, David T. Thomas, and Qiao Ping Xiang. ...
'' * ''
Alangium rotundifolium ''Alangium rotundifolium'' is a tree in the family Cornaceae. The specific epithet ' is from the Latin meaning "rounded leaves". Description ''Alangium rotundifolium'' grows up to tall with a trunk diameter of up to . The smooth to scaly bark i ...
'' * ''
Alangium salviifolium ''Alangium salviifolium'', commonly known as sage-leaved alangium, is a flowering plant in the Cornaceae family. It is also commonly known as Ankolam in Malayalam, Ankola in Kannada, Akola or Ankol in Hindi and Alanji in Tamil. In India, Its mos ...
'' ** subsp. ''hexapetalum'' * ''
Alangium scandens ''Alangium'' is a small genus of flowering plants. The genus is included either in a broad view of the dogwood family Cornaceae, or as the sole member of its own family Alangiaceae.Qiu-Yun (Jenny) Xiang, David T. Thomas, and Qiao Ping Xiang. ...
'' * ''
Alangium sinicum ''Alangium'' is a small genus of flowering plants. The genus is included either in a broad view of the dogwood family Cornaceae, or as the sole member of its own family Alangiaceae.Qiu-Yun (Jenny) Xiang, David T. Thomas, and Qiao Ping Xiang. ...
'' * '' Alangium taiwanianum'' * ''
Alangium tetrandrum ''Alangium'' is a small genus of flowering plants. The genus is included either in a broad view of the dogwood family Cornaceae, or as the sole member of its own family Alangiaceae.Qiu-Yun (Jenny) Xiang, David T. Thomas, and Qiao Ping Xia ...
'' * ''
Alangium tonkinense ''Alangium'' is a small genus of flowering plants. The genus is included either in a broad view of the dogwood family Cornaceae, or as the sole member of its own family Alangiaceae.Qiu-Yun (Jenny) Xiang, David T. Thomas, and Qiao Ping Xiang. ...
'' * ''
Alangium uniloculare ''Alangium'' is a small genus of flowering plants. The genus is included either in a broad view of the dogwood family Cornaceae, or as the sole member of its own family Alangiaceae.Qiu-Yun (Jenny) Xiang, David T. Thomas, and Qiao Ping Xiang. ...
'' (syn. '' Alangium griffithii'') * ''
Alangium villosum ''Alangium villosum'' is a species of flowering plant in the family Cornaceae, native to Java. Subtaxa It was previously considered to contain ten described and one undescribed subspecies, but these were split out to their own species, such as ...
'' ** subsp. ''bussyanum'' ** subsp. ''ferrugineum'' ** var. ''javanicum'' ** subsp. ''parviflorum'' ** subsp. ''pilosum'' ** subsp. ''polyosmoides'' ** subsp. ''tomentosum'' ** subsp. ''vitiense'' ** subsp. ''warburgianum'' * ''
Alangium yunnanense ''Alangium'' is a small genus of flowering plants. The genus is included either in a broad view of the dogwood family Cornaceae, or as the sole member of its own family Alangiaceae.Qiu-Yun (Jenny) Xiang, David T. Thomas, and Qiao Ping Xiang. ...
''


Characteristics

Differences from the other
genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...
in Cornaceae include articulated
pedicel Pedicle or pedicel may refer to: Human anatomy *Pedicle of vertebral arch, the segment between the transverse process and the vertebral body, and is often used as a radiographic marker and entry point in vertebroplasty and kyphoplasty procedures ...
s,
subulate The following is a list of terms which are used to describe leaf morphology in the description and taxonomy of plants. Leaves may be simple (a single leaf blade or lamina) or compound (with several leaflets). The edge of the leaf may be regular o ...
bract In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves. They may be smaller, larger, or of ...
s,
bitegmic In biology, an integument is the tissue surrounding an organism's body or an organ within, such as skin, a husk, shell, germ or rind. Etymology The term is derived from ''integumentum'', which is Latin for "a covering". In a transferred, or ...
seeds and the single-seeded fruit. The entire or lobed
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
are alternate. The
bisexual Bisexuality is a romantic or sexual attraction or behavior toward both males and females, or to more than one gender. It may also be defined to include romantic or sexual attraction to people regardless of their sex or gender identity, whi ...
(rarely unisexual)
nectar Nectar is a sugar-rich liquid produced by plants in glands called nectaries or nectarines, either within the flowers with which it attracts pollinating animals, or by extrafloral nectaries, which provide a nutrient source to animal mutualists ...
iferous flowers are arranged in axillary
cymes An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed on ...
. The flowers have 4-10 small sepals and 4-10 linear
petal Petals are modified Leaf, leaves that surround the reproductive parts of flowers. They are often advertising coloration, brightly colored or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''c ...
s. There are 4–40
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...
s distributed in a single
cycle Cycle, cycles, or cyclic may refer to: Anthropology and social sciences * Cyclic history, a theory of history * Cyclical theory, a theory of American political history associated with Arthur Schlesinger, Sr. * Social cycle, various cycles in soc ...
. The
ovary The ovary is an organ in the female reproductive system that produces an ovum. When released, this travels down the fallopian tube into the uterus, where it may become fertilized by a sperm. There is an ovary () found on each side of the body. ...
is
inferior Inferior may refer to: * Inferiority complex * An Anatomical terms of location#Superior and inferior, anatomical term of location * Inferior angle of the scapula, in the human skeleton *Inferior (book), ''Inferior'' (book), by Angela Saini * ''The ...
and
bilocular A locule (plural locules) or loculus (plural loculi) (meaning "little place" in Latin) is a small cavity or compartment within an organ or part of an organism (animal, plant, or fungus). In angiosperms (flowering plants), the term ''locule'' usu ...
(sometimes unilocular). The fruit is
drupe In botany, a drupe (or stone fruit) is an indehiscent fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pit'', ''stone'', or '' pyrena'') of hardened endocarp with a seed (''kernel'') ...
. A detailed description of ''Alangium'' can be found at
Flora of China (journal) ''Flora of China'' is a scientific publication aimed at describing the plants native to China. The project is a collaborative scientific effort to publish the first modern English-language account of the 31,000 species of vascular plants of C ...
.Haining Qin and Chamlong Phengklai. 2007. ''Alangium'' pages 304-308. In: Wu Zhengyi, Peter H. Raven, and Hong Deyuan (editors). 1994 onward. ''Flora of China vol. 13: Clusiaceae - Araliaceae.'' Science Press: Beijing, China; and Missouri Botanical Garden Press: St. Louis, MO, USA. (vol. 13) (set). (See ''External links'' below). Detailed botanical illustrations are available for several species.Wu Zhengyi, Peter H. Raven, Hong Deyuan (editors), and Zhang Libing (illustrations editor). 2008. ''Flora of China Illustrations'', vol. 13: Clusiaceae - Araliaceae: 328-333. Science Press: Beijing, China; and Missouri Botanical Garden Press: St. Louis, MO, USA. .


Taxonomy

In 2011, a phylogenetic analysis of
DNA sequence DNA sequencing is the process of determining the nucleic acid sequence – the order of nucleotides in DNA. It includes any method or technology that is used to determine the order of the four bases: adenine, guanine, cytosine, and thymine. Th ...
s showed that ''Alangium'' is sister to '' Cornus''. Since 1939, ''Alangium'' has been divided into four
sections Section, Sectioning or Sectioned may refer to: Arts, entertainment and media * Section (music), a complete, but not independent, musical idea * Section (typography), a subdivision, especially of a chapter, in books and documents ** Section sig ...
: Conostigma, Rhytidandra, Marlea, and Alangium. Some authors have raised Marlea and Rhytidandra to generic rank. The intergeneric
classification Classification is a process related to categorization, the process in which ideas and objects are recognized, differentiated and understood. Classification is the grouping of related facts into classes. It may also refer to: Business, organizat ...
of ''Alangium'' will require a few changes.


Traditional uses

One species, ''Alangium chinense'' (), is considered one of the fifty fundamental herbs in traditional Chinese medicine.


References


External links


''Alangium''
A
Index Nominum Genericorum
At

At

At
Research and Collections
At
Smithsonian National Museum of Natural History

''Alangium''
In: Volume 1
Of: Encyclopédie méthodique: botanique
At: Biodiversity Heritage Library

List of lower taxa''Alangium''.pdf''Alangium''Chinese Plant NamesAlangiaceaeFOC vol. 13Family ListFlora of ChinaeFloras

''Alangium''
At:Plant Names
At:IPNI

CRC World Dictionary of Plant Names: A-C
At:Botany & Plant Science
At:Life Science
At:CRC PressGermplasm Resources Information Network: ''Alangium''
*
Flora of Pakistan: ''Alangium''Flora of Madagascar: ''Alangium''Flora of China checklist: ''Alangium''
{{Authority control Plants used in traditional Chinese medicine Cornales genera