Agrimonia Procera
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''Agrimonia procera'' is a species of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
belonging to the family
Rosaceae Rosaceae (), the rose family, is a medium-sized family of flowering plants that includes 4,828 known species in 91 genera. The name is derived from the type genus ''Rosa''. Among the most species-rich genera are ''Alchemilla'' (270), ''Sorbus ...
. It is native to Europe and Southern Africa.


Description

A
herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition of t ...
perennial plant A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
which grows to about 1 m tall, with a branched or unbranched green stem covered in
glandular In animals, a gland is a group of cells in an animal's body that synthesizes substances (such as hormones) for release into the bloodstream (endocrine gland) or into cavities inside the body or its outer surface (exocrine gland). Structure De ...
hairs which are 2 mm long or slightly longer. The leaves are
pinnate Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and in ...
with 3-6 pairs of oval, distinctly serrated primary leaflets, an irregular number of secondary leaflets, and a similar or slightly larger terminal leaflet. The underside of the leaves are studded with yellow, shiny glands which produce a sweet aroma. Flowers are arranged in long
raceme A raceme ( or ) or racemoid is an unbranched, indeterminate type of inflorescence bearing flowers having short floral stalks along the shoots that bear the flowers. The oldest flowers grow close to the base and new flowers are produced as the s ...
s at the top of the stem(s). Each flower has 5 yellow
petal Petals are modified Leaf, leaves that surround the reproductive parts of flowers. They are often advertising coloration, brightly colored or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''c ...
s, 5-20
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...
s and 2
carpels Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) ''pistils'' ...
. The
hypanthium In angiosperms, a hypanthium or floral cup is a structure where basal portions of the calyx, the corolla, and the stamens form a cup-shaped tube. It is sometimes called a floral tube, a term that is also used for corolla tube and calyx tube. It ...
develops into a characteristic brown, oval or bell-shaped fruit with deep lateral grooves and a double ring of hooked bristles around its centre, which contains 1 or 2
achene An achene (; ), also sometimes called akene and occasionally achenium or achenocarp, is a type of simple dry fruit produced by many species of flowering plants. Achenes are monocarpellate (formed from one carpel) and indehiscent (they do not ope ...
s. It is easy to confuse fragrant agrimony with common agrimony, which has a similar distribution and habitat. The key differences are that fragrant agrimony has longer hairs (and no short hairs) on the stem, leaves that are more deeply and acutely serrated and with more glands on the underside, and fruits with shorter grooves and reflexed bristles. These hooked bristles are the most useful feature to use when it is not possible to compare the two species side-by-side: if you imagine they emerge from the fruit's "equator", then most of the bristles point "southwards" or at most straight out in common agrimony, but in fragrant agrimony some distinctly point "northwards" (towards the stalk). left, A mature fruit of fragrant agrimony


Ecology

The habitat of fragrant agrimony in Britain and Ireland is woodland margin, scrub, road verges and tall grassland, on circumneutral (neither acid nor alkaline), sometimes damp, soils. It is confined to the lowlands, being known only up to 335 m at Fortingall, in Perthshire. It is a bigger, bushier and more leafy plant than common agrimony and is far less common, although its overall range is similar. It is most abundant along rides in
oak An oak is a tree or shrub in the genus ''Quercus'' (; Latin "oak tree") of the beech family, Fagaceae. There are approximately 500 extant species of oaks. The common name "oak" also appears in the names of species in related genera, notably ''L ...
or sometimes
ash Ash or ashes are the solid remnants of fires. Specifically, ''ash'' refers to all non-aqueous, non- gaseous residues that remain after something burns. In analytical chemistry, to analyse the mineral and metal content of chemical samples, ash ...
woods, or in field edges nearby. It flowers rather later than common agrimony, typically from late July in southern England. Little is known about the interactions between fragrant agrimony and other species. The British database of insects and their food plants lists only one aphid, ''
Acyrthosiphon ''Acyrthosiphon'' is a genus of aphids belonging to the family Aphididae The Aphididae are a very large insect family in the aphid superfamily ( Aphidoidea), of the order Hemiptera. These insects suck the sap from plant leaves. Several thousan ...
malvae'' (Mosley, 1841) that has been observed on this plant. The hooked seeds are ideal for facilitating dispersal by animals, but which are primarily involved is not known.


Distribution

The native distribution of fragrant agrimony is thought to include almost all of
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
and
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...
, but none of the territory between. This strikingly
disjunct distribution In biology, a taxon with a disjunct distribution is one that has two or more groups that are related but considerably separated from each other geographically. The causes are varied and might demonstrate either the expansion or contraction of a s ...
is highly unusual and could be due to transportation of the seeds by migrating wild birds or a more recent introduction by humans. In South Africa, however, it is considered native and is officially categorised as "least concern", although some sources, such as the citizen science website iNaturalist, describe it as invasive there.


Uses

Fragrant agrimony is palatable to livestock and occurs in grazed woodland, meadows and pastures. Like other species of agrimony, it is reported to have beneficial effects on the health of experimental animals and to reduce inflammation in cells ''in vitro''. Chemicals under investigation include the bitter-tasting compound agrimoniin and various flavonoids. Despite having a bitter taste, agrimony is popular in
herbal tea Herbal teas, also known as herbal infusions and less commonly called tisanes (UK and US , US also ), are beverages made from the infusion or decoction of herbs, spices, or other plant material in hot water. Oftentimes herb tea, or the plain term ...
s and is often touted as having health benefits in humans, although medical sources stress that evidence about efficacy and safety is generally lacking. Claims about its use in the past may be exaggerated: the Roman author
Celsus Celsus (; grc-x-hellen, Κέλσος, ''Kélsos''; ) was a 2nd-century Greek philosopher and opponent of early Christianity. His literary work, ''The True Word'' (also ''Account'', ''Doctrine'' or ''Discourse''; Greek: grc-x-hellen, ΛόγΠ...
, who was influential in medicine for over a thousand years, only mentions agrimony as a possible cure for snake bites. Most advertisements for herbal teas specify common rather than fragrant agrimony, but it is not always clear that people have differentiated between the two.


References


External links

{{Taxonbar, from=Q953646
procera Procera is a hypothetical clade of amphibians that includes salamanders and caecilians but not frogs. A close relationship between salamanders and caecilians is a competing hypothesis to the more widely supported view that salamanders and frogs a ...
Taxa named by Karl Friedrich Wilhelm Wallroth