Agriades Glandon
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''Agriades glandon'', the Arctic blue or Glandon blue, is a species of butterfly in the family Lycaenidae. It in found in Eurasia and
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
.


Range

In North America, this species is found from Alaska east to
Newfoundland Newfoundland and Labrador (; french: Terre-Neuve-et-Labrador; frequently abbreviated as NL) is the easternmost province of Canada, in the country's Atlantic region. The province comprises the island of Newfoundland and the continental region ...
, south through the mountains to Washington, northern Arizona, and northern New Mexico. In Europe, it is found in mountainous areas like the Pyrenees and Alps, as well as the far north. It is also found in parts of Russia, including Siberia, and Kamchatka. Its habitats include arctic tundra, subarctic and subalpine forests, mountain meadows, and bogs.


Description

The wingspan is 17–26 mm. The male wing uppersides are silver-coloured, steely blue or pale shining blue and become increasingly brown towards the edges. The female wing uppersides are almost entirely brown but with a slightly bluish pollination in the basal region. The incidence of darkening increases with altitude. All wings usually have small, dark disk spots that are sometimes surrounded by white. Both sexes have white-rimmed black dots on the pale grey-brown ground of the forewing underside, while the underside of the hind wings reveals striking white spots and occasionally orange-colored dots on the greyish-brown ground. Despite a large geographic variability in the pattern of the wing, the species is, in Europe, clearly distinguishable from other bluish species in a complex of three species widely separated geographically. Only two species overlap in range in Europe that could be confused. In the Alps and the Pyrenees, the more local and less frequent, '' Aricia nicias'', has fine underside markings and a white stripe. In the Pyrenees, ''
Agriades pyrenaicus ''Agriades pyrenaicus'', the Gavarnie blue, is a Palearctic butterfly of the family Lycaenidae. It is found in the Asturias mountains of north-western Spain, the Pyrenees, the southern Balkan Peninsula, Turkey, the Caucasus and Armenia. The habit ...
'' may also fly in the same locations. In this species the underside forewing submarginal spots are bold black not faint grey-brown and the black markings on the underside hindwing are relatively greatly reduced or absent. Other similar species do not overlap in distribution. They are '' Agriades zullichi'' (Spain) and ''
Agriades aquilo ''Agriades glandon'', the Arctic blue or Glandon blue, is a species of butterfly in the family Lycaenidae. It in found in Eurasia and North America. Range In North America, this species is found from Alaska east to Newfoundland, south through th ...
'' (Arctic Europe), and are sometimes viewed as subspecies of ''Agriades glandon''.


Ecology

The butterfly flies from mid-May to September depending on the location. Clarke 2022 lists ''
Astralagus alpinus ''Astragalus'' is a large genus of over 3,000 species of herbs and small shrubs, belonging to the legume family Fabaceae and the subfamily Faboideae. It is the largest genus of plants in terms of described species. The genus is native to temp ...
'', '' Saxifraga aizoides'' and '' Saxifraga oppositifolia'' as larval food plants in the European part of its range for ''Agriades aquilo''. For ''Agriades glandon'', Clarke lists '' Androsace'' species '' A. alpina'', A. chamaejasme'', '' A. lactea'', '' A. laggeri'', '' A. obtusifolia'', '' A. villosa'' and '' A. vitaliana''; '' Soldanella alpina'' and ''
Soldanella pusilla The genus ''Soldanella'', commonly known in English as snowbell, includes about 15 species of flowering plants native to European mountains, from the Pyrenees, the Apennines, the Alps, the Carpathians and the Balkans. They grow in woods, damp p ...
''; and '' Oxytropis campestris''. Further recorded food plants for ''Agriades glandon'' include ''
Androsace bungeana ''Androsace'', commonly known as rock jasmine, is a genus of flowering plants in the family Primulaceae, second only to '' Primula'' in the number of species. It is predominantly Arctic–alpine, with many species in the Himalayas (where the ge ...
'' and '' Androsace septentrionalis'', '' Diapensia lapponica'', '' Vaccinium'', and '' Saxifraga'' species ''
Saxifraga bronchialis ''Saxifraga'' is the largest genus in the family Saxifragaceae, containing about 465 species of holarctic perennial plants, known as saxifrages or rockfoils. The Latin word ''saxifraga'' means literally "stone-breaker", from Latin ' ("rock" or " ...
'' and '' Saxifraga spinulosa''.


Subspecies and taxonomy

Several
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
have been described: *''A. g. glandon'' (de Prunner, 1798) (Pyrenees, Alps) *''A. g. aquilo'' (Boisduval, 1832) (Arctic Europe and Arctic Canada) *''A. g. franklinii'' (Curtis, 1835) (Alaska) *''A. g. wosnesenskii'' (Ménétriés, 1855) (Srednesibirskoe, northeastern Siberia, Kamchatka) has the pale spots of underside smaller and more widely separated from one another, especially on the hindwing. *''A. g. rustica'' (Edwards, 1865) (Colorado) *''A. g. aquilina'' (Staudinger, 1901) (polar tundra of Siberia) *''A. g. megalo'' (McDunnough, 1927) (British Columbia) *''A. g. bryanti'' (Leussler, 1935) (Alaska, Northwest Territories) *''A. g. lacustris'' (Freeman, 1939) (Manitoba) *''A. g. centrohelvetica'' Rezbanyai-Reser, 1981 (Switzerland) *''A. g. punctatus'' Austin, 1998 (Arizona) *''A. g. cassiope'' Emmel & Emmel, 1998 (California) *''A. g. kelsoni'' Emmel & Emmel, 1998 (California) *''A. g. ustjuzhanini'' Yakivlev & Churkin, 2003 (Mongolia) *''A. g. saluki'' Churkin, 2005 *''A. g. brutus'' Churkin, 2005 (eastern Sayan Mountains) *''A. g. rubini'' Churkin, 2005 (eastern Kazakhstan) *''A. g. batchimeg'' Churkin, 2005 (Mongolia) *''A. g. labrador'' Schmidt, Scott & Kondla, 2006 (Labrador, Newfoundland) The relationships between these taxa are not yet fully understood. Many authors treat ''A. g. aquilo'' as a separate species. Some authors also consider all the above North American subspecies to belong to one separate species, which they call ''Agriades franklinii''. Subspecies ''rustica'' and ''cassiope'' are sometimes also treated as valid species.


References


External links

*
Arctic blue
Butterflies and Moths of North America

{{Taxonbar, from=Q22104603, from2=Q1266072 Agriades Butterflies of North America Butterflies of Europe Insects of the Arctic Butterflies described in 1798