:Common names: ''cantil, Mexican cantil, Mexican ground pit viper,
[ Gloyd HK, Conant R. 1990. ''Snakes of the Agkistrodon Complex: A Monographic Review''. Society for the Study of Amphibians and Reptiles. 614 pp. 52 plates. LCCN 89-50342. .] cantil viper,
[Mehrtens JM. 1987. ''Living Snakes of the World in Color''. New York: Sterling Publishers. 480 pp. .] black moccasin,
[Campbell JA, Lamar WW. 2004. ''The Venomous Reptiles of the Western Hemisphere''. Comstock Publishing Associates, Ithaca and London. 870 pp. 1500 plates. .] Mexican moccasin,
more.''
''Agkistrodon bilineatus'' is a highly
venomous
Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a ...
pit viper species
In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...
found in
Mexico
Mexico ( Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guate ...
and
Central America
Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
as far south as
Honduras.
Description
These are heavy-bodied snakes, and share the same general body structure with
copperheads. They average around in length and have a broad, triangular-shaped head with small
eye
Eyes are organs of the visual system. They provide living organisms with vision, the ability to receive and process visual detail, as well as enabling several photo response functions that are independent of vision. Eyes detect light and conv ...
s that have vertical
pupil
The pupil is a black hole located in the center of the iris of the eye that allows light to strike the retina.Cassin, B. and Solomon, S. (1990) ''Dictionary of Eye Terminology''. Gainesville, Florida: Triad Publishing Company. It appears black ...
s.
Coloration can vary, but most are brown or black, with darker brown or black banding, sometimes with
white
White is the lightness, lightest color and is achromatic (having no hue). It is the color of objects such as snow, chalk, and milk, and is the opposite of black. White objects fully diffuse reflection, reflect and scattering, scatter all the ...
or cream-colored accents. ''A. taylori'' is known for being more elaborately patterned, often having distinct
tan-colored banding, sometimes with
orange
Orange most often refers to:
*Orange (fruit), the fruit of the tree species '' Citrus'' × ''sinensis''
** Orange blossom, its fragrant flower
* Orange (colour), from the color of an orange, occurs between red and yellow in the visible spectrum ...
or
yellow
Yellow is the color between green and orange on the spectrum of light. It is evoked by light with a dominant wavelength of roughly 575585 nm. It is a primary color in subtractive color systems, used in painting or color printing. In ...
accents that can almost appear gold in color. There are the following distinctive yellow and/or white lines on the head: a vertical line on the rostral and mental, a fine line on the canthus continuing above and beyond the eye to the neck, a broader line on the upper lip from the anterior nasal to the last labial. Juveniles are almost always distinctly banded, with bright
green
Green is the color between cyan and yellow on the visible spectrum. It is evoked by light which has a dominant wavelength of roughly 495570 Nanometre, nm. In subtractive color systems, used in painting and color printing, it is created by ...
or yellow tail tips, which they use to lure prey. As they age, their pattern and coloration fade and darken.
Common names
Mexican ground pit viper, cantil viper,
cantil,
Mexican moccasin,
neotropical moccasin,
[Gotch AF. 1986. ''Reptiles – Their Latin Names Explained''. Poole, UK: Blandford Press. 176 pp. .] Mexican yellow-lipped viper.
[Parker HW, Grandison AGC. 1977. ''Snakes – a natural history. Second Edition''. British Museum (Natural History) and Cornell University Press. 108 pp. 16 plates. LCCCN 76-54625. (cloth), (paper).]
The common name, "cantil", is based on the
Tzeltal Tzeltal may refer to:
* Tzeltal people, an ethnic group of Mexico
* Tzeltal language
Tzeltal or Tseltal () is a Mayan language spoken in the Mexican state of Chiapas, mostly in the municipalities of Ocosingo, Altamirano, Huixtán, Tenejapa, ...
word ''kantiil'' "yellow lips."
Geographic range
Mexico and Central America. On the Pacific side it is found from southern
Sonora
Sonora (), officially Estado Libre y Soberano de Sonora ( en, Free and Sovereign State of Sonora), is one of the 31 states which, along with Mexico City, comprise the Federal Entities of Mexico. The state is divided into 72 municipalities; the ...
in Mexico south through
Guatemala,
El Salvador
El Salvador (; , meaning " The Saviour"), officially the Republic of El Salvador ( es, República de El Salvador), is a country in Central America. It is bordered on the northeast by Honduras, on the northwest by Guatemala, and on the south by ...
, and
Honduras. The
type locality
Type locality may refer to:
* Type locality (biology)
* Type locality (geology)
See also
* Local (disambiguation)
* Locality (disambiguation)
{{disambiguation ...
given is "Pacific coast of Guatemala."
Conservation status
This species is classified as Near Threatened (NT) on the
IUCN Red List of Threatened Species
The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biolog ...
(v3.1, 2001).
A species is listed as such when it has been evaluated against the criteria but does not qualify for Critically Endangered, Endangered or Vulnerable now, but is close to qualifying for, or is likely to qualify for a threatened category in the near future. The population trend is down. Year assessed: 2007.
The primary ecological concern is habitat loss.
Behavior
Much like the
cottonmouth, with whom it shares a genus, this species has a reputation for having a nasty disposition and being extremely dangerous, a reputation probably not well deserved. They are generally shy by nature, and if threatened their first instinct is to rely on camouflage. If unable to do so they will use a threat display to ward off potential predators. The tightly coiled animal will raise the last several inches of its tail, this portion often being bright yellow or green in juveniles and a faded yellow or green in adults, the animal will then
tail vibrate, creating a loud whipping sound against its coils or surroundings. This particular behavior is very reminiscent of caudal luring, though in a more violent fashion and is often accompanied by a strike or less commonly a gaping display similar to that of ''A. piscivorus''. They generally will only display these behaviors when given no other choice. In captivity, ''A. bilineatus'' are often known for aggression stemming from their characteristic lack of predictability.
Reproduction
Breeding occurs in the spring, and like most other viper species, cantils are
ovoviviparous
Ovoviviparity, ovovivipary, ovivipary, or aplacental viviparity is a term used as a "bridging" form of reproduction between egg-laying oviparous and live-bearing viviparous reproduction. Ovoviviparous animals possess embryos that develop insi ...
, giving birth to 5–20 young at a time.
Captivity
Export from Mexico is not permitted, but cantils of both species are often captive-bred, making them frequently available in the exotic pet trade. They are also well represented in
zoo
A zoo (short for zoological garden; also called an animal park or menagerie) is a facility in which animals are kept within enclosures for public exhibition and often bred for conservation purposes.
The term ''zoological garden'' refers to zoo ...
s throughout North America and Europe.
Venom
According to Gloyd and Conant (1990), "this species is greatly feared throughout its range," in some areas even more so than ''
Bothrops asper
Terciopelo (''Bothrops asper'') is a species of pit vipers occurring at low to moderate elevations in northeast Mexico and Central America, and into South America where it is known from elevations up to 2600 meters in the Colombian and Ecuador ...
''. In
Sonora
Sonora (), officially Estado Libre y Soberano de Sonora ( en, Free and Sovereign State of Sonora), is one of the 31 states which, along with Mexico City, comprise the Federal Entities of Mexico. The state is divided into 72 municipalities; the ...
, Mexico, it is feared more than any other reptile. In
Nicaragua
Nicaragua (; ), officially the Republic of Nicaragua (), is the largest Sovereign state, country in Central America, bordered by Honduras to the north, the Caribbean Sea, Caribbean to the east, Costa Rica to the south, and the Pacific Ocean to ...
, it is considered the country's most dangerous snake.
[Warrell DA. 2004. Snakebites in Central and South America: Epidemiology, Clinical Features, and Clinical Management. ''in'' .]
Bite symptoms, in general, may include nothing more than local pain, swelling and discoloration, but those from adult specimens can cause massive swelling and
necrosis
Necrosis () is a form of cell injury which results in the premature death of cells in living tissue by autolysis. Necrosis is caused by factors external to the cell or tissue, such as infection, or trauma which result in the unregulated dig ...
. Campbell and Lamar (1989) suggested that, due to the necrosis,
amputation
Amputation is the removal of a limb by trauma, medical illness, or surgery. As a surgical measure, it is used to control pain or a disease process in the affected limb, such as malignancy or gangrene. In some cases, it is carried out on ind ...
may be required in one out of every six cases. Some bites were fatal within only a few hours. Gaige (1936) cites one case in which a woman in Motul,
Yucatán
Yucatán (, also , , ; yua, Yúukatan ), officially the Free and Sovereign State of Yucatán,; yua, link=no, Xóot' Noj Lu'umil Yúukatan. is one of the 31 states which comprise the federal entities of Mexico. It comprises 106 separate mun ...
, Mexico was bitten by a specimen and died within a few hours. Alvarez del Toro (1983) reports
gangrenous
Gangrene is a type of tissue death caused by a lack of blood supply. Symptoms may include a change in skin color to red or black, numbness, swelling, pain, skin breakdown, and coolness. The feet and hands are most commonly affected. If the gan ...
tissue falling away in fragments, eventually to expose the underlying bones, describing this is as "spontaneous amputation" of the necrotic wound.
In
Honduras, Cruz (1987) describes the bite symptoms as being similar to those of ''
Bothrops'' species, although more severe considering the small size of these snakes. They include immediate and severe pain, oozing of blood from the fang punctures, considerable
edema
Edema, also spelled oedema, and also known as fluid retention, dropsy, hydropsy and swelling, is the build-up of fluid in the body's tissue. Most commonly, the legs or arms are affected. Symptoms may include skin which feels tight, the area ma ...
,
nosebleed
A nosebleed, also known as epistaxis, is bleeding from the nose. Blood can flow down into the stomach, and cause nausea and vomiting. In more severe cases, blood may come out of both nostrils. Rarely, bleeding may be so significant that low blo ...
, bleeding of the gums, marked
hematuria
Hematuria or haematuria is defined as the presence of blood or red blood cells in the urine. “Gross hematuria” occurs when urine appears red, brown, or tea-colored due to the presence of blood. Hematuria may also be subtle and only detectable ...
, general
petechia
A petechia () is a small red or purple spot (≤4 mm in diameter) that can appear on the skin, conjunctiva, retina, and mucous membranes which is caused by haemorrhage of capillaries. The word is derived from Italian , 'freckle,' of obscure origin ...
e,
shock
Shock may refer to:
Common uses Collective noun
*Shock, a historic commercial term for a group of 60, see English numerals#Special names
* Stook, or shock of grain, stacked sheaves
Healthcare
* Shock (circulatory), circulatory medical emerge ...
,
kidney failure
Kidney failure, also known as end-stage kidney disease, is a medical condition in which the kidneys can no longer adequately filter waste products from the blood, functioning at less than 15% of normal levels. Kidney failure is classified as eit ...
and local
necrosis
Necrosis () is a form of cell injury which results in the premature death of cells in living tissue by autolysis. Necrosis is caused by factors external to the cell or tissue, such as infection, or trauma which result in the unregulated dig ...
.
Polyvalent Antivenom, produced by the Instituto Clodomiro Picado
Taxonomy
A new subspecies, ''A. b. lemosespinali'', was described by
H. M. Smith &
Chiszar (2001) based on a single specimen from near Palma Sola,
Veracruz
Veracruz (), formally Veracruz de Ignacio de la Llave (), officially the Free and Sovereign State of Veracruz de Ignacio de la Llave ( es, Estado Libre y Soberano de Veracruz de Ignacio de la Llave), is one of the 31 states which, along with Me ...
, Mexico.
[ Smith H. M., Chiszar D.. 2001. A new subspecies of cantil (''Agkistrodon bilineatus'') from Central Veracruz, Mexico (Reptilia: Serpentes). Bulletin of the Maryland Herpetological Society, 37: 130-136.]''
Agkistrodon howardgloydi'', ''
Agkistrodon russeolus'' and ''
Agkistrodon taylori'' were formerly considered subspecies.
References
External links
*
*
''Agkistrodon bilineatus''a
VN Illustrated Database of Mexican Biodiversity Accessed 26 November 2006.
{{Taxonbar, from=Q1926277
bilineatus
Reptiles of Mexico
Reptiles of Guatemala
Reptiles of Honduras
Snakes of Central America
Taxa named by Albert Günther
Reptiles described in 1863