Agama Lizard
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''Agama'' (from
Sranan Tongo Sranan Tongo (also Sranantongo "Surinamese tongue," Sranan, Surinaams, Surinamese, Surinamese Creole) is an English-based creole language that is spoken as a ''lingua franca'' by approximately 550,000 people in Suriname. Developed originally amo ...
meaning "lizard") is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of small-to-moderate-sized, long-tailed, insectivorous Old World lizards. The genus ''Agama'' includes at least 37
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
in
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
, especially
sub-Saharan Africa Sub-Saharan Africa is, geographically, the area and regions of the continent of Africa that lies south of the Sahara. These include West Africa, East Africa, Central Africa, and Southern Africa. Geopolitically, in addition to the List of sov ...
, where most regions are home to at least one species.
Eurasia Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelago a ...
n
agamids Agamidae is a family of over 300 species of iguanian lizards indigenous to Africa, Asia, Australia, and a few in Southern Europe. Many species are commonly called dragons or dragon lizards. Overview Phylogenetically, they may be sister to the I ...
are largely assigned to genus ''
Laudakia ''Laudakia'' is a genus of lizards, commonly known as Asian rock agamas, in the family Agamidae. The genus is found mostly in Asia, with some species in Southern Europe. Taxonomy Some species of ''Laudakia'', ''sensu lato'', are now recognized i ...
''. The various species differ in size, ranging from about in length, when fully grown. Their colour also differs between species, between genders, and according to mood; for example, a dominant male in display mode is far brighter than when it has been caught, beaten by another male, or otherwise alarmed. Females tend to be less colourful than the males of the species. According to species, agamas live in forest, in bush, among rocks and on crags, but where their habitat has been cleared, or simply occupied by humans, some species also adapt to life in villages and compounds, for example inside the thatch of huts and other sheltering crevices. Agamids' hind legs generally are long and powerful; and the lizards can run and leap swiftly when alarmed. Agamas are diurnal, active during the day. They can tolerate higher temperatures than most reptiles, but when temperatures approach 38 °C (100 °F) they generally shelter in the shade. Males frequently threaten each other by nodding, weaving, and displaying their brightest colours to establish dominance. If that is insufficient, they lash their tails and threaten each other with open jaws. The jaws are very powerful, and older males commonly have damaged tails as souvenirs of past combat. Females may sometimes chase and fight one another, and hatchlings mimic the adults' behaviour. Agamas are mainly insectivorous, hunting prey by sight and snatching it opportunistically. Their incisor-like front teeth and powerful jaws are adapted to dealing with quite large, hard prey. They also may eat eggs of other lizards, and sometimes feed on vegetable matter, such as suitable grass, berries, and seeds. Though not formally polygamous, dominant males commonly accommodate several females at a time in their territory. During courtship, and also when asserting his territory, the male bobs his head in display; this gives rise to some of the common names, such as
Afrikaans Afrikaans (, ) is a West Germanic language that evolved in the Dutch Cape Colony from the Dutch vernacular of Holland proper (i.e., the Hollandic dialect) used by Dutch, French, and German settlers and their enslaved people. Afrikaans gra ...
''koggelmander'' (literally, "little mocking man"). Females occasionally initiate courtship by offering their hindquarters to the male and provoking him to catch her. Typically the breeding season is timed for eggs to be laid during the season after the rains. Eggs are laid in clutches of up to 12, depending on species and the size of the female.


Etymology and taxonomy

In the ''
10th edition of Systema Naturae The 10th edition of ''Systema Naturae'' is a book written by Swedish naturalist Carl Linnaeus and published in two volumes in 1758 and 1759, which marks the starting point of zoological nomenclature. In it, Linnaeus introduced binomial nomencla ...
'' of 1758, Linnaeus used the name Agama (pg. 288) as the species ''Lacerta Agama'' (with Agama originally capitalized to indicate a name in apposition rather than a Latin adjective, which he would have made lowercase). His own earlier description from 1749 was derived from Seba, who described and illustrated a number of lizards as ''Salamandra amphibia'' and ''Salamandra Americana'', said to resemble in some ways a chameleon lizard and that supposedly came (in error) from "America." Seba did not use the term "agama", however. Linnaeus repeated Seba's error in stating that the lizards lived in the Americas habitat in America" and he included other types of lizards shown and mentioned by Seba under his species name ''Agama''. Daudin later created the new genus, ''Agama'', to incorporate various African and Asian lizards, as well as species from Mexico, the Caribbean, Central America, and South America. He noted that the name ''agama'' was used by inhabitants of Guiana for a species that he included in the genus ''Agama''. The word "agama" has been traced to West African
Gbe languages The Gbe languages (pronounced ) form a cluster of about twenty related languages stretching across the area between eastern Ghana and western Nigeria. The total number of speakers of Gbe languages is between four and eight million. The most widel ...
as a name for the
chameleon Chameleons or chamaeleons (family Chamaeleonidae) are a distinctive and highly specialized clade of Old World lizards with 202 species described as of June 2015. The members of this family are best known for their distinct range of colors, bein ...
. The word was brought to Dutch Guiana (modern
Suriname Suriname (; srn, Sranankondre or ), officially the Republic of Suriname ( nl, Republiek Suriname , srn, Ripolik fu Sranan), is a country on the northeastern Atlantic coast of South America. It is bordered by the Atlantic Ocean to the north ...
) by imported West African slaves and was then used in local creole languages for types of local lizards. Linnaeus may have taken the name "agama" from some unidentified source in the mistaken belief that the reptiles came from the Americas as indicated by Seba. The name "agama" has no connection to either Greek ''agamos'' "unmarried" (as a supposed Latin feminine ''agama'') or to Greek ''agamai'' "wonder" as sometimes suggested. Because of the confusion over the actual taxon that was the basis for the name ''Agama agama'', Wagner, et al. (2009) designated a neotype (ZFMK 15222), using a previously described specimen from Cameroon in the collection of the Zoologisches Forschungsmuseum Alexander Koenig in Bonn.


Species

Listed alphabetically. ''
Nota bene (, or ; plural form ) is a Latin phrase meaning "note well". It is often abbreviated as NB, n.b., or with the ligature and first appeared in English writing . In Modern English, it is used, particularly in legal papers, to draw the atten ...
'': A
binomial authority In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
in parentheses indicates that the species was originally described in a genus other than ''Agama''.


References


Further reading

* Manthey, Ulrich; Schuster, Norbert (1996). ''Agamid Lizards''. U.S.A.: T.F.H Publications Inc. 189 pp. . * Spawls, Stephen; Howell, Kim M.; Drewes, Robert C. (2006). ''Reptiles and Amphibians of East Africa''. Princeton, New Jersey: Princeton University Press. .


External links


Information on Agamids in captivity
* {{Taxonbar, from=Q1153386 Agamid lizards of Africa Lizard genera Taxa named by François Marie Daudin