''Afrovenator'' (; "African hunter") is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

megalosaurid Megalosauridae is a monophyletic family of carnivorous theropod dinosaurs within the group Megalosauroidea. Appearing in the Middle Jurassic, megalosaurids were among the first major radiation of large theropod dinosaurs. They were a relative ...

theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

from the

Middle Jurassic The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations co ...

Period of northern

Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...

Discovery and naming

The remains of ''Afrovenator'' were discovered in 1993 in the

Tiourarén Formation The Tiourarén Formation is a geological Formation (geology), formation in the Agadez Region of Niger whose strata were originally thought to be Early Cretaceous. However, re-interpretation of the sediments showed that they are probably Middle Jur ...

of the department of

Agadez Agadez ( Air Tamajeq: ⴰⴶⴰⴷⴰⵣ, ''Agadaz''), formerly spelled Agadès, is the fifth largest city in Niger, with a population of 110,497 based on the 2012 census. The capital of Agadez Region, it lies in the Sahara desert, and is also ...

Niger ) , official_languages = , languages_type = National languagesHauterivian The Hauterivian is, in the geologic timescale, an age in the Early Cretaceous Epoch or a stage in the Lower Cretaceous Series. It spans the time between 132.9 ± 2 Ma and 129.4 ± 1.5 Ma (million years ago). The Hauterivian is preceded by the Va ...

Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 129.4 ± 1.5 Ma (million years ago) and 121.4 ± 1.0 Ma). It is a subdivision of the Early Cretaceous Epoch (or Lower Cretaceous Series). It is preceded ...

stages of the early

Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...

Period, or approximately 132 to 125 million years ago (Sereno et al. 1994). However, re-interpretation of the sediments showed that they are probably Mid-Jurassic in age, dating ''Afrovenator'' to the

Bathonian In the geologic timescale the Bathonian is an age and stage of the Middle Jurassic. It lasted from approximately 168.3 Ma to around 166.1 Ma (million years ago). The Bathonian Age succeeds the Bajocian Age and precedes the Callovian Age. Strat ...

to Oxfordian stages, between 167 and 161 mya. The

sauropod Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their bo ...

Jobaria ''Jobaria'' is a genus of sauropod dinosaur that lived in what is now Niger during the middle Jurassic Period, between 164 and 161 million years ago. ''Jobaria'' is currently the only known valid sauropod from the Tiouraren, where it was discov ...

'', whose remains were first mentioned in the same paper which named ''Afrovenator'', is also known from this

formation Formation may refer to: Linguistics * Back-formation, the process of creating a new lexeme by removing or affixes * Word formation, the creation of a new word by adding affixes Mathematics and science * Cave formation or speleothem, a secondar ...

. ''Afrovenator'' is known from a single relatively complete skeleton,

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...

UC OBA 1, featuring most of the

skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, the ...

minus its top (likewise the

mandible In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower tooth, teeth in place. The mandible sits beneath the maxilla. It is the only movabl ...

, or lower jaws, are lacking apart from the prearticular bone), parts of the

spinal column The vertebral column, also known as the backbone or spine, is part of the axial skeleton. The vertebral column is the defining characteristic of a vertebrate in which the notochord (a flexible rod of uniform composition) found in all chordates ...

, partial forelimbs, a partial

pelvis The pelvis (plural pelves or pelvises) is the lower part of the trunk, between the abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also called bony pelvis, or pelvic skeleton). The ...

, and most of the hind limbs. This skeleton is housed at the

University of Chicago The University of Chicago (UChicago, Chicago, U of C, or UChi) is a private research university in Chicago, Illinois. Its main campus is located in Chicago's Hyde Park neighborhood. The University of Chicago is consistently ranked among the b ...

. The generic name comes from the

Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...

''afer'', "African", and ''venator'', "hunter". There is one named

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

, ''Afrovenator abakensis''. The generic name refers to its predatory nature, and its provenance from Africa. The specific name refers to Abaka, the

Tuareg The Tuareg people (; also spelled Twareg or Touareg; endonym: ''Imuhaɣ/Imušaɣ/Imašeɣăn/Imajeɣăn'') are a large Berber ethnic group that principally inhabit the Sahara in a vast area stretching from far southwestern Libya to southern A ...

name for the region of Niger where the

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

was found. The original short description of both genus and species is found in a 1994 paper which appeared in the prestigious journal ''

Science Science is a systematic endeavor that builds and organizes knowledge in the form of testable explanations and predictions about the universe. Science may be as old as the human species, and some of the earliest archeological evidence for ...

''. The primary author was well-known

American American(s) may refer to: * American, something of, from, or related to the United States of America, commonly known as the "United States" or "America" ** Americans, citizens and nationals of the United States of America ** American ancestry, pe ...

paleontologist Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fossi ...

Paul Sereno Paul Callistus Sereno (born October 11, 1957) is a professor of paleontology at the University of Chicago and a National Geographic "explorer-in-residence" who has discovered several new dinosaur species on several continents, including at sites ...

, with Jeffrey Wilson, Hans Larsson, Didier Dutheil, and

Hans-Dieter Sues Hans-Dieter Sues (born January 13, 1956) is a German-born American paleontologist who is Senior Scientist and Curator of Vertebrate Paleontology at the National Museum of Natural History of the Smithsonian Institution in Washington, DC. He receiv ...

as coauthors.

Description

Afrovenatorinae_Size_Comparison_by_PaleoGeek

Judging from the one skeleton known, this dinosaur was about 8 meters (26 feet) long, from snout to tail tip, and had a weight of about 1 tonne according to

Gregory S. Paul Gregory Scott Paul (born December 24, 1954) is an American freelance researcher, author and illustrator who works in paleontology, and more recently has examined sociology and theology. He is best known for his work and research on theropod dino ...

. Thomas R. Holtz Jr. estimated it at 7.6 meters (25 feet) in length and 453–907 kg (1,000–2,000 lbs) in weight. In 2016 it was given a lower estimation of 6.8 meters (22.3 feet) in length, 1.9 meters (6.2 feet) tall at the hips and 790 kg (1.742 lbs) in weight. Sereno stressed that the general build was gracile and that the forelimbs and the lower leg were relatively long: the

humerus The humerus (; ) is a long bone in the arm that runs from the shoulder to the elbow. It connects the scapula and the two bones of the lower arm, the radius and ulna, and consists of three sections. The humeral upper extremity consists of a roun ...

has length of forty centimetres and the tibia and fourth metatarsal measure 687 and 321 millimetres respectively, as compared to a thighbone length of seventy-six centimetres. Several

autapomorphies In phylogenetics, an autapomorphy is a distinctive feature, known as a derived trait, that is unique to a given taxon. That is, it is found only in one taxon, but not found in any others or outgroup taxa, not even those most closely related to t ...

have been established, traits that distinguish ''Afrovenator'' from its nearest relatives. The depression in which the

antorbital fenestra An antorbital fenestra (plural: fenestrae) is an opening in the skull that is in front of the eye sockets. This skull character is largely associated with archosauriforms, first appearing during the Triassic Period. Among extant archosaurs, birds ...

is located, has a front end in the form of a lobe. The third neck vertebra has a low rectangular spine. The crescent-shaped wrist bone is very flat. The first metacarpal has a broadly expanding contact surface with the second metacarpal. The foot of the

pubic bone In vertebrates, the pubic region ( la, pubis) is the most forward-facing (ventral and anterior) of the three main regions making up the coxal bone. The left and right pubic regions are each made up of three sections, a superior ramus, inferior ra ...

is notched from behind. In general the skull is rather flat, its height being less than three times its length, which cannot be exactly determined because the

praemaxilla The premaxilla (or praemaxilla) is one of a pair of small cranial bones at the very tip of the upper jaw of many animals, usually, but not always, bearing teeth. In humans, they are fused with the maxilla. The "premaxilla" of therian mammal has ...

e are lacking. The

maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The t ...

, which has a long front branch, bears fourteen teeth, as can be deduced from the tooth sockets: the teeth themselves have been lost. There is a small maxillary fenestra, which does not reach the edge of the antorbital depression and is located behind a promaxillary fenestra. The

lacrimal bone The lacrimal bone is a small and fragile bone of the facial skeleton; it is roughly the size of the little fingernail. It is situated at the front part of the medial wall of the orbit. It has two surfaces and four borders. Several bony landmarks of ...

has a distinctive rounded horn on top. The lower branch of the

postorbital bone The ''postorbital'' is one of the bones in vertebrate skulls which forms a portion of the dermal skull roof and, sometimes, a ring about the orbit. Generally, it is located behind the postfrontal and posteriorly to the orbital fenestra. In some ve ...

is transversely wide. The

jugal bone The jugal is a skull bone found in most reptiles, amphibians and birds. In mammals, the jugal is often called the malar or zygomatic. It is connected to the quadratojugal and maxilla, as well as other bones, which may vary by species. Anatomy ...

is short, deep and pneumatised.

Classification

Most analyses place ''Afrovenator'' within the

Megalosauridae Megalosauridae is a monophyletic family of carnivorous theropod dinosaurs within the group Megalosauroidea. Appearing in the Middle Jurassic, megalosaurids were among the first major radiation of large theropod dinosaurs. They were a relatively ...

, which was formerly a "wastebasket family" which contained many large and hard-to-classify theropods, but has since been redefined in a meaningful way, as a

sister taxon In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and t ...

to the family

Spinosauridae The Spinosauridae (or spinosaurids) are a clade or family of tetanuran theropod dinosaurs comprising ten to seventeen known genera. They came into prominence during the Cretaceous period. Spinosaurid fossils have been recovered worldwide, includin ...

within the

Megalosauroidea Megalosauroidea (meaning 'great/big lizard forms') is a superfamily (or clade) of tetanuran theropod dinosaurs that lived from the Middle Jurassic to the Late Cretaceous period. The group is defined as ''Megalosaurus bucklandii'' and all taxa sha ...

. A 2002 analysis, focused mainly on the noasaurids, found ''Afrovenator'' to be a basal megalosaurid. However, it did not include ''

Dubreuillosaurus ''Dubreuillosaurus'' is a genus of carnivorous dinosaur from the middle Jurassic Period. It is a megalosaurid theropod. Its fossils were found in France. The only named species, ''Dubreuillosaurus valesdunensis'', was originally described as a s ...

'' (formerly ''

Poekilopleuron ''Poekilopleuron'' (meaning "varied ribs") is a genus of tetanuran dinosaur, which lived during the middle Bathonian of the Jurassic, about 168 to 166 million years ago. The genus has been used under many different spelling variants, although on ...

valesdunensis''), which could affect the results in that region of the

cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ...

(Carrano ''et al''. 2002). Other more recent and more complete cladistic analyses show ''Afrovenator'' in a group of Megalosauridae with ''

Eustreptospondylus ''Eustreptospondylus'' ( ; meaning "true ''Streptospondylus''") is a genus of megalosaurid Theropoda, theropod dinosaur, from the Oxfordian stage of the Late Jurassic period (some time between 163 and 154 million years ago) in southern Eng ...

'' and ''Dubreuillosaurus''. This group is either called Megalosaurinae (Allain 2002) or

Eustreptospondylinae Megalosauridae is a monophyletic family of carnivorous theropod dinosaurs within the group Megalosauroidea. Appearing in the Middle Jurassic, megalosaurids were among the first major radiation of large theropod dinosaurs. They were a relative ...

(Holtz ''et al''. 2004). The latter study also includes ''

Piatnitzkysaurus ''Piatnitzkysaurus'' ( meaning "Piatnitzky's lizard") is a genus of megalosauroid theropod dinosaur that lived approximately 179 to 177 million years ago during the lower part of the Jurassic Period in what is now Argentina. ''Piatnitzkysaurus'' ...

'' in this taxon. A study by Matthew Carrano from 2012 placed ''Afrovenator'' in a megalosaurid

Afrovenatorinae Megalosauridae is a monophyletic family of carnivorous theropod dinosaurs within the group Megalosauroidea. Appearing in the Middle Jurassic, megalosaurids were among the first major radiation of large theropod dinosaurs. They were a relatively ...

. A few alternative hypotheses have been presented for ''Afrovenators relationships. In Sereno's original description, ''Afrovenator'' was found to be a basal spinosauroid (he at the time used the name " Torvosauroidea"), outside of Spinosauridae and Megalosauridae (which he called " Torvosauridae"). Finally, another recent study places ''Afrovenator'' outside of the Megalosauroidea completely, and instead finds it more closely related to ''

Allosaurus ''Allosaurus'' () is a genus of large carnosaurian theropod dinosaur that lived 155 to 145 million years ago during the Late Jurassic epoch (Kimmeridgian to late Tithonian). The name "''Allosaurus''" means "different lizard" alluding to ...

'' (Rauhut 2003). This is the only study to draw this conclusion. Afrovenator mount (1)

The

phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spec ...

according to Carrano ''et al.'' (2012) is shown by this cladogram.

Discovery and naming

Description

Classification

References

Further reading