Afroboletus Luteolus
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''Afroboletus'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
fungi A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from ...
in the family
Boletaceae The Boletaceae are a family of mushroom-forming fungi, primarily characterised by small pores on the spore-bearing hymenial surface (at the underside of the mushroom), instead of gills as are found in most agarics. Nearly as widely distributed ...
. The genus, circumscribed in 1981, contains seven species found in
tropical Africa Although tropical Africa is mostly familiar to the West for its rainforests, this biogeographic realm of Africa is far more diverse. While the tropics are thought of as regions with hot moist climates, which are caused by latitude and the tropi ...
.


Taxonomy

When defining the genus, Pegler and Young placed it in the family Strobilomycetaceae, and considered ''Afroboletus'' to be a "primitive" and possibly ancestral member of the group because of the form of its
basidia A basidium () is a microscopic sporangium (a spore-producing structure) found on the hymenophore of fruiting bodies of basidiomycete fungi which are also called tertiary mycelium, developed from secondary mycelium. Tertiary mycelium is highly-c ...
and
spore In biology, a spore is a unit of sexual or asexual reproduction that may be adapted for dispersal and for survival, often for extended periods of time, in unfavourable conditions. Spores form part of the life cycles of many plants, algae, f ...
s, the reticulation of the
stem Stem or STEM may refer to: Plant structures * Plant stem, a plant's aboveground axis, made of vascular tissue, off which leaves and flowers hang * Stipe (botany), a stalk to support some other structure * Stipe (mycology), the stem of a mushro ...
, tropical distribution, and non-
mycorrhiza   A mycorrhiza (from Greek μύκης ', "fungus", and ῥίζα ', "root"; pl. mycorrhizae, mycorrhiza or mycorrhizas) is a symbiotic association between a fungus and a plant. The term mycorrhiza refers to the role of the fungus in the plant ...
l requirements of its species.


Description

Fruit bodies The sporocarp (also known as fruiting body, fruit body or fruitbody) of fungi is a multicellular structure on which spore-producing structures, such as basidia or asci, are borne. The fruitbody is part of the sexual phase of a fungal life cyc ...
of ''Afroboletus'' species have fleshy
caps Caps are flat headgear. Caps or CAPS may also refer to: Science and technology Computing * CESG Assisted Products Service, provided by the U.K. Government Communications Headquarters * Composite Application Platform Suite, by Java Caps, a Java ...
that are hemispherical or convex to applanate (horizontally flattened). As it ages, the cap surface becomes fuliginous (sooty) and black, developing pustules or scales. The cap margin is appendiculate, meaning that
partial veil In mycology, a partial veil (also called an inner veil, to differentiate it from the "outer", or universal veil) is a temporary structure of tissue found on the fruiting bodies of some basidiomycete fungi, typically agarics. Its role is to isolate ...
remnants hang along the cap margin. On the cap underside, the pore surface comprises tubes that are adnately attached to the stipe. The pore surface appears swollen, and is initially white before turning greyish or pale pinkish. The tubes are relatively long, measuring , and there are typically 1–2 pores per millimeter. Stipes are slender, cylindrical, firm, and solid (i.e., not hollow). The colour of the stipe surface is similar to that of the cap, and also eventually turns blackish. The
veil A veil is an article of clothing or hanging cloth that is intended to cover some part of the head or face, or an object of some significance. Veiling has a long history in European, Asian, and African societies. The practice has been prominent ...
is persistent, fluffy, and sometimes leaves a
ring Ring may refer to: * Ring (jewellery), a round band, usually made of metal, worn as ornamental jewelry * To make a sound with a bell, and the sound made by a bell :(hence) to initiate a telephone connection Arts, entertainment and media Film and ...
on the stipe. The
flesh Flesh is any aggregation of soft tissues of an organism. Various multicellular organisms have soft tissues that may be called "flesh". In mammals, including humans, ''flesh'' encompasses muscle Skeletal muscles (commonly referred to as mu ...
is whitish, discoloring black on exposure.
Spore print 300px, Making a spore print of the mushroom ''Volvariella volvacea'' shown in composite: (photo lower half) mushroom cap laid on white and dark paper; (photo upper half) cap removed after 24 hours showing pinkish-tan spore print. A 3.5-centimeter ...
s are dark
fuscous {{Short pages monitor