Aeshna Isosceles
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''Aeshna isoceles'' is a small hawker
dragonfly A dragonfly is a flying insect belonging to the infraorder Anisoptera below the order Odonata. About 3,000 extant species of true dragonfly are known. Most are tropical, with fewer species in temperate regions. Loss of wetland habitat threate ...
that is found in Europe, mostly around the Mediterranean, and the lowlands of North Africa. Its common name in English is green-eyed hawker. In Britain it is a rare and local species and is known as the Norfolk hawker. It has a brown colour with green eyes and clear wings and also a yellow triangular mark on the second
abdominal The abdomen (colloquially called the belly, tummy, midriff, tucky or stomach) is the part of the body between the thorax (chest) and pelvis, in humans and in other vertebrates. The abdomen is the front part of the abdominal segment of the torso. ...
segment which gave rise to its
scientific name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
. It used to be in the genus ''
Anaciaeschna ''Anaciaeschna'' is a genus of dragonflies in the family Aeshnidae. Species of ''Anaciaeschna'' are found in Africa, Asia and Australia. ''Anaciaeschna'' are large, tawny brown dragonflies. They are nomadic and crepuscular. Species The genus '' ...
'' as it has several differences from the other members of the genus ''
Aeshna ''Aeshna'', or the mosaic darners, is a genus of dragonflies from the family Aeshnidae. Species within this genus are generally known as "hawkers" (Old World) or "darners" (New World). Description These are relatively large dragonflies. T ...
''. Its specific name is often spelt ''isosceles''.


Identification

''A. isoceles'' is one of only two brown hawkers found in Europe, the other is '' A. grandis''. Both have a brown thorax and abdomen but ''A. isoceles'' has green eyes and clear wings and a diagnostic yellow triangular mark on the second
abdominal The abdomen (colloquially called the belly, tummy, midriff, tucky or stomach) is the part of the body between the thorax (chest) and pelvis, in humans and in other vertebrates. The abdomen is the front part of the abdominal segment of the torso. ...
segment. The hindwings have an amber patch at their base. In contrast ''A. grandis'' has yellowish wings and blueish eyes. The green eye of ''A. isoceles'' stands out even in flight and in practice it is not difficult to tell these two dragonflies apart. In addition to the morphological differences ''A. isoceles'' is on the wing much earlier in the year than '' A. grandis''.


Distribution and habitat

''A. isoceles'' is found in central Europe and around the Mediterranean and, the lowlands of North Africa. It is more common in eastern Europe than the south western Europe; it occurs in Spain and Portugal but is local. It is found in wet areas, ponds, ditches and marshes, with dense vegetation and, in studies carried out in England, was found to be associated with Water-soldier (''Stratiotes aloides'').


Status in Britain

The Norfolk hawker has always been a scarce and local insect in Britain. It used to be found in the Cambridgeshire fens but by the early 1980s the populations had greatly declined. It is now confined to relatively unpolluted fens and grazing marshes in the Broadlands of Norfolk and north-east
Suffolk Suffolk () is a ceremonial county of England in East Anglia. It borders Norfolk to the north, Cambridgeshire to the west and Essex to the south; the North Sea lies to the east. The county town is Ipswich; other important towns include Lowes ...
. It can be found in
Hickling Broad Hickling Broad is a nature reserve 4 km south-east of Stalham, north-east of Norwich in Norfolk. It is managed by the Norfolk Wildlife Trust. It is a National Nature Reserve and part of the Upper Thurne Broads and Marshes Site of Special ...
and two national nature reserves: Mid-Yare NNR and
Ludham - Potter Heigham NNR Ludham - Potter Heigham Marshes is a biological Site of Special Scientific Interest north-east of Norwich in Norfolk. Part of the site is a Nature Conservation Review site, Grade I, and a National Nature Reserve. It is part of the Broadland Ram ...
and at Castle Marshes in the
Barnby Broad and Marshes Barnby Broad and Marshes is a Site of Special Scientific Interest (SSSI) in the Waveney district of the English county of Suffolk. The site is in size. It is in the parishes of North Cove and Barnby, located between Beccles and Lowestoft ...
SSSI. Since 2011 the species has also been recorded in the Stour valley in east Kent where egg laying has been observed and it appears to be spreading. It is protected under Schedule 5 of the Wildlife and Countryside Act 1981 and listed in Category 1 ( endangered) in the British Red Data Books on Insects.


Behaviour

It is one of the earliest ''
Aeshna ''Aeshna'', or the mosaic darners, is a genus of dragonflies from the family Aeshnidae. Species within this genus are generally known as "hawkers" (Old World) or "darners" (New World). Description These are relatively large dragonflies. T ...
'' dragonflies to be on the wing with a flight period from May to August. Adults do not spend as much time on the wing as other ''Aeshnas''. Males will fly around over a stretch of water defending a territory and if the pond is small the male will hover over the centre of the pond. Unlike other aeshnas, where the adults seem to be continuously on the wing beating up and down their territory, male ''A. isoceles'' come to rest on vegetation from time to time. Females oviposit onto plants and the eggs hatch in about 2 weeks. Larval development takes 2 years.


Systematics

This species was first described as ''Libellula quadrifasciata, var. 36. isoceles'' by Muller in 1764. It has since been called ''Aeshna rufescens'' and ''Aeshna chysophthalmus'' and more recently ''Anaciaeschna isoceles''. It is by this last name that it is referred to in many books. It has since been included into the genus ''
Aeshna ''Aeshna'', or the mosaic darners, is a genus of dragonflies from the family Aeshnidae. Species within this genus are generally known as "hawkers" (Old World) or "darners" (New World). Description These are relatively large dragonflies. T ...
'' and in many books is called ''Aeshna isoceles'': however the original specific name was ''isosceles''. Dijkstra and Lewington (2006) and Boudot JP., ''et al.'' (2009) both call it ''Aeshna isoceles'' whereas Askew, R.R. (2004),and earlier books, refer to it as ''Aeshna isosceles''.


Notes


References

* Askew, R.R. (2004) ''The Dragonflies of Europe''. (revised ed.) Harley Books. * d'Aguilar, J., Dommanget, JL., and Prechac, R. (1986) ''A field guide to the Dragonflies of Britain, Europe and North Africa''. Collins. pp336. * Boudot JP., ''et al.'' (2009) ''Atlas of the Odonata of the Mediterranean and North Africa''. ''Libellula'' Supplement 9:1-256. * Dijkstra, K-D.B & Lewington, R. (2006) ''Field Guide to the Dragonflies of Britain and Europe''. British Wildlife Publishing. . *


External links

* {{Taxonbar, from=Q1435436 Aeshnidae Dragonflies of Europe Norfolk Broads Insects described in 1767 Taxa named by Otto Friedrich Müller