Actinostrobus Pyramidalis - Flickr - Kevin Thiele
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''Actinostrobus'' is a genus of
conifer Conifers are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single ...
ous
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
s in the
Cupressaceae Cupressaceae is a conifer family, the cypress family, with worldwide distribution. The family includes 27–30 genera (17 monotypic), which include the junipers and redwoods, with about 130–140 species in total. They are monoecious, subdio ...
(cypress family). Common names include cypress, sandplain-cypress and cypress-pine, the last of these shared by the closely related genus ''
Callitris ''Callitris'' is a genus of coniferous trees in the Cupressaceae (cypress family). There are 16 recognized species in the genus, of which 13 are native to Australia and the other three (''C. neocaledonica, C. sulcata'' and ''C. p ...
''.


Species

There are three species in the genus, all
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to southwestern
Western Australia Western Australia (commonly abbreviated as WA) is a state of Australia occupying the western percent of the land area of Australia excluding external territories. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to th ...
: A 2010 study of ''Actinostrobus'' and ''Callitris'' has placed all three species of ''Actinostrobus'' within an expanded ''Callitris'' based on analysis of 42 morphological and anatomical characters. They are
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
s or small
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
s, reaching tall. The
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
are
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, which ...
, of two forms; juvenile needle-like leaves long on young seedlings (but occasional into adulthood in ''A. acuminatus''), and scale-like adult leaves, long with only the apex free. The leaves are arranged in six rows along the twigs, in alternating whorls of three. The male
cones A cone is a three-dimensional geometric shape that tapers smoothly from a flat base (frequently, though not necessarily, circular) to a point called the apex or vertex. A cone is formed by a set of line segments, half-lines, or lines conn ...
are small, long, and are located at the tips of the twigs. The female cones start out similarly inconspicuous, maturing in eighteen to twenty months to long and wide, globular to acute-ovoid, with six thick, woody scales, arranged in two whorls of three, and a further nine to fifteen thin, sterile basal scales. The cones remain closed on the trees for many years, opening only after being scorched by a
bushfire A wildfire, forest fire, bushfire, wildland fire or rural fire is an unplanned, uncontrolled and unpredictable fire in an area of combustible vegetation. Depending on the type of vegetation present, a wildfire may be more specifically identif ...
; this then releases the
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...
s to grow on the newly cleared burnt ground. The closest relative of ''Actinostrobus'' is ''
Callitris ''Callitris'' is a genus of coniferous trees in the Cupressaceae (cypress family). There are 16 recognized species in the genus, of which 13 are native to Australia and the other three (''C. neocaledonica, C. sulcata'' and ''C. p ...
'', which is much more widespread, occurring in most of
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
, and differs in its cones lacking the basal whorls of small sterile scales. The wood of ''Actinostrobus'' is light, soft and aromatic, but the plants are too small for any significant use. They are occasionally planted as ornamental shrubs, but their use is restricted by the high risks imposed by their very high flammability in bushfires.


References

* *Farjon, A. (2005). ''Monograph of Cupressaceae and Sciadopitys''. Royal Botanic Gardens, Kew. .
Gymnosperm Database - ''Actinostrobus''
{{Taxonbar, from=Q1472074 Flora of Western Australia Pinales of Australia Conifer genera