Acraea Lycoa
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''Acraea lycoa'', the lycoa acraea, is a
butterfly Butterflies are insects in the macrolepidopteran clade Rhopalocera from the Order (biology), order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The ...
in the family
Nymphalidae The Nymphalidae are the largest family of butterflies, with more than 6,000 species distributed throughout most of the world. Belonging to the superfamily Papilionoidea, they are usually medium-sized to large butterflies. Most species have a red ...
which is native to the African tropics and subtropics.


Region

It is found in
Sierra Leone Sierra Leone,)]. officially the Republic of Sierra Leone, is a country on the southwest coast of West Africa. It is bordered by Liberia to the southeast and Guinea surrounds the northern half of the nation. Covering a total area of , Sierra ...
,
Liberia Liberia (), officially the Republic of Liberia, is a country on the West African coast. It is bordered by Sierra Leone to Liberia–Sierra Leone border, its northwest, Guinea to its north, Ivory Coast to its east, and the Atlantic Ocean ...
,
Ivory Coast Ivory Coast, also known as Côte d'Ivoire, officially the Republic of Côte d'Ivoire, is a country on the southern coast of West Africa. Its capital is Yamoussoukro, in the centre of the country, while its largest city and economic centre is ...
,
Ghana Ghana (; tw, Gaana, ee, Gana), officially the Republic of Ghana, is a country in West Africa. It abuts the Gulf of Guinea and the Atlantic Ocean to the south, sharing borders with Ivory Coast in the west, Burkina Faso in the north, and To ...
,
Togo Togo (), officially the Togolese Republic (french: République togolaise), is a country in West Africa. It is bordered by Ghana to the west, Benin to the east and Burkina Faso to the north. It extends south to the Gulf of Guinea, where its c ...
,
Benin Benin ( , ; french: Bénin , ff, Benen), officially the Republic of Benin (french: République du Bénin), and formerly Dahomey, is a country in West Africa. It is bordered by Togo to the west, Nigeria to the east, Burkina Faso to the north ...
,
Nigeria Nigeria ( ), , ig, Naìjíríyà, yo, Nàìjíríà, pcm, Naijá , ff, Naajeeriya, kcg, Naijeriya officially the Federal Republic of Nigeria, is a country in West Africa. It is situated between the Sahel to the north and the Gulf o ...
,
Cameroon Cameroon (; french: Cameroun, ff, Kamerun), officially the Republic of Cameroon (french: République du Cameroun, links=no), is a country in west-central Africa. It is bordered by Nigeria to the west and north; Chad to the northeast; the C ...
,
Bioko Bioko (; historically Fernando Po; bvb, Ëtulá Ëria) is an island off the west coast of Africa and the northernmost part of Equatorial Guinea. Its population was 335,048 at the 2015 census and it covers an area of . The island is located of ...
,
Angola , national_anthem = " Angola Avante"() , image_map = , map_caption = , capital = Luanda , religion = , religion_year = 2020 , religion_ref = , coordina ...
, the
Democratic Republic of the Congo The Democratic Republic of the Congo (french: République démocratique du Congo (RDC), colloquially "La RDC" ), informally Congo-Kinshasa, DR Congo, the DRC, the DROC, or the Congo, and formerly and also colloquially Zaire, is a country in ...
,
Sudan Sudan ( or ; ar, السودان, as-Sūdān, officially the Republic of the Sudan ( ar, جمهورية السودان, link=no, Jumhūriyyat as-Sūdān), is a country in Northeast Africa. It shares borders with the Central African Republic t ...
,
Uganda }), is a landlocked country in East Africa East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territor ...
,
Rwanda Rwanda (; rw, u Rwanda ), officially the Republic of Rwanda, is a landlocked country in the Great Rift Valley of Central Africa, where the African Great Lakes region and Southeast Africa converge. Located a few degrees south of the Equator ...
,
Burundi Burundi (, ), officially the Republic of Burundi ( rn, Repuburika y’Uburundi ; Swahili language, Swahili: ''Jamuhuri ya Burundi''; French language, French: ''République du Burundi'' ), is a landlocked country in the Great Rift Valley at the ...
,
Ethiopia Ethiopia, , om, Itiyoophiyaa, so, Itoobiya, ti, ኢትዮጵያ, Ítiyop'iya, aa, Itiyoppiya officially the Federal Democratic Republic of Ethiopia, is a landlocked country in the Horn of Africa. It shares borders with Eritrea to the ...
,
Kenya ) , national_anthem = "Ee Mungu Nguvu Yetu"() , image_map = , map_caption = , image_map2 = , capital = Nairobi , coordinates = , largest_city = Nairobi , ...
,
Tanzania Tanzania (; ), officially the United Republic of Tanzania ( sw, Jamhuri ya Muungano wa Tanzania), is a country in East Africa within the African Great Lakes region. It borders Uganda to the north; Kenya to the northeast; Comoro Islands and ...
and
Zambia Zambia (), officially the Republic of Zambia, is a landlocked country at the crossroads of Central Africa, Central, Southern Africa, Southern and East Africa, although it is typically referred to as being in Southern Africa at its most cent ...
.


Description

''A. lycoa'' is a very variable species nearly allied to '' johnstoni''; it differs from this in having the light basal or median area of the hindwing distally rounded or occasionally entirely absent. The fore wing has normally 5-7 white or whitish spots, in the male sometimes dull and very indistinct; the first is placed in 1b close to the distal margin and is quite free and rounded or partly joined to the spot in 2, but never extends so far basad as the latter; the third is placed in 4 near to the distal margin quite free or touching the spot in 5; the spots in 5 and 6 are of equal length, joined to one another and to small spots in cellules 8 and 9; the cell always unicolorous without spots. * ''lycoa'' Godt. both wings thinly scaled, semi-transparent; spots of the forewing indistinct; basal area of the hindwing large, dull reddish yellow, distally rather sharply defined against the grey marginal band. female: wings thinly scaled, dark grey with distinct white markings; marginal band media, of the hindwing broad, not sharply defined.
Sierra Leone Sierra Leone,)]. officially the Republic of Sierra Leone, is a country on the southwest coast of West Africa. It is bordered by Liberia to the southeast and Guinea surrounds the northern half of the nation. Covering a total area of , Sierra ...
to
Nigeria Nigeria ( ), , ig, Naìjíríyà, yo, Nàìjíríà, pcm, Naijá , ff, Naajeeriya, kcg, Naijeriya officially the Federal Republic of Nigeria, is a country in West Africa. It is situated between the Sahel to the north and the Gulf o ...
. * ''media'' Eltr. (57 a, as ''lycoa''). male. Spots of the forewing more distinct; hindwing in the middle somewhat tinged with reddish yellow but without distinct basal area. female with darker ground-colour and sharply defined marginal band on the hindwing.
Cameroon Cameroon (; french: Cameroun, ff, Kamerun), officially the Republic of Cameroon (french: République du Cameroun, links=no), is a country in west-central Africa. It is bordered by Nigeria to the west and north; Chad to the northeast; the C ...
s to Toro sub-region, Toro. * ''bukoba'' Eltr. male Forewing dark olive-brown; spots light yellow, sharply defined; basal area of the hindwing ochre-yellow. female similar to the male, but darker, with white or whitish markings. The spot in 1b of the forewing in both sexes entirely free and broadly separated from the spot in 2.
Urundi Burundi (, ), officially the Republic of Burundi ( rn, Repuburika y’Uburundi ; Swahili: ''Jamuhuri ya Burundi''; French: ''République du Burundi'' ), is a landlocked country in the Great Rift Valley at the junction between the African Gre ...
. * ''entebbia'' Eltr. The spot in 1b of the forewing quite free; ground-colour in the male yellowish grey; spots of the forewing small, dull yellowish; basal area of the hind wing small, not reaching the inner margin; ground-colour in the female very dark, spots of the forewing white, basal area of the hindwing whitish yellow.
Uganda }), is a landlocked country in East Africa East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territor ...
. * ''tirika'' Eltr. Spot 1b of the forewing widely separated; ground-colour of the fore wing in the olive-brown, in the female very dark, spots small; basal area of the hind wing yellowish.
British East Africa East Africa Protectorate (also known as British East Africa) was an area in the African Great Lakes occupying roughly the same terrain as present-day Kenya from the Indian Ocean inland to the border with Uganda in the west. Controlled by Britai ...
. * ''fallax'' Rogenh. (57 c). Ground-colour in both sexes nearly black; spots of the forewing small, in the male yellowish, in the female white, spot 1b widely separated from spot 2; basal area of the hindwing sharply defined, yellowish.
German East Africa German East Africa (GEA; german: Deutsch-Ostafrika) was a German colony in the African Great Lakes region, which included present-day Burundi, Rwanda, the Tanzania mainland, and the Kionga Triangle, a small region later incorporated into Mozam ...
:
Kilimandjaro Mount Kilimanjaro () is a dormant volcano in Tanzania. It has three volcanic cones: Kibo, Mawenzi, and Shira. It is the highest mountain in Africa and the highest free-standing mountain above sea level in the world: above sea level and ab ...
. * ''kenia'' Eltr. Smaller than the other races; ground-colour nearly black; spots of the forewing in the male lemon-yellow, in the female white; basal area of the hindwing in both sexes lemon-yellow. British East Africa: Kenia;
Kikuyu Kikuyu or Gikuyu (Gĩkũyũ) mostly refers to an ethnic group in Kenya or its associated language. It may also refer to: * Kikuyu people, a majority ethnic group in Kenya *Kikuyu language, the language of Kikuyu people *Kikuyu, Kenya, a town in Cent ...
. * ''aequalis'' Rothsch. and Jord. Sexes similar; spots of the forewing and basal area of the hindwing dull yellowish; marginal band of the hindwing yellowish dark brown.
Abyssinia The Ethiopian Empire (), also formerly known by the exonym Abyssinia, or just simply known as Ethiopia (; Amharic and Tigrinya: ኢትዮጵያ , , Oromo: Itoophiyaa, Somali: Itoobiya, Afar: ''Itiyoophiyaa''), was an empire that historica ...
.


Biology

The habitat consists of forests. The larvae feed on ''
Theobroma cacao ''Theobroma cacao'', also called the cacao tree and the cocoa tree, is a small ( tall) evergreen tree in the family Malvaceae. Its seeds, cocoa beans, are used to make chocolate liquor, cocoa solids, cocoa butter and chocolate. The largest pro ...
'', '' Pouzolzia guineensis'', '' Pouzolzia parasitica'', ''
Aneilema ''Aneilema'' is a genus of monocotyledonous plants of approximately 60 species. The vast majority of the species are native to sub-Saharan Africa, but a few are found in Oceania and one, '' Aneilema brasiliense'', is from South America. It is the ...
'' and '' Fleurya'' species.


Taxonomy

It is a member of the ''
Acraea jodutta ''Acraea jodutta'', the jodutta acraea, is a butterfly in the family Nymphalidae. It is found in Guinea, Sierra Leone, Liberia, Ivory Coast, Ghana, Togo, Nigeria, Cameroon, Equatorial Guinea, São Tomé and Príncipe, Gabon, the Republic of the ...
''
species group In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...
– but see also Pierre & Bernaud, 2014 Pierre & Bernau, 2014 Classification et Liste Synonymique des Taxons du Genre ''Acraea'
pdf
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References


External links


''Die Gross-Schmetterlinge der Erde'' 13: Die Afrikanischen Tagfalter. Plate XIII 57
''c'' and also ''c'' as ''fallax''
Images representing ''Acraea lycoa''
at
Bold In typography, emphasis is the strengthening of words in a text with a font in a different style from the rest of the text, to highlight them. It is the equivalent of prosody stress in speech. Methods and use The most common methods in W ...
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''Acraea lycoa lycoa''
at Pteron Butterflies described in 1819 lycoa Butterflies of Africa Taxa named by Jean-Baptiste Godart {{Heliconiinae-stub