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''Achyranthes arborescens'' (common names - Chaff tree, Soft-wood) is a plant in the
Amaranthaceae Amaranthaceae is a family of flowering plants commonly known as the amaranth family, in reference to its type genus ''Amaranthus''. It includes the former goosefoot family Chenopodiaceae and contains about 165 genera and 2,040 species, making it ...
family endemic to
Norfolk Island Norfolk Island (, ; Norfuk: ''Norf'k Ailen'') is an external territory of Australia located in the Pacific Ocean between New Zealand and New Caledonia, directly east of Australia's Evans Head and about from Lord Howe Island. Together with ...
. It is a critically endangered species under the Australian Federal government's
Environment Protection and Biodiversity Conservation Act The ''Environment Protection and Biodiversity Conservation Act 1999'' (Cth) is an Act of the Parliament of Australia that provides a framework for protection of the Australian environment, including its biodiversity and its natural and cultu ...
.


Description

''A. arborescens'' is a soft-wooded tree growing up to a height of 9 m. Its leaves are on slender stalks which are 5–15 mm long and have hairs lying close to them (appressed hairs). The leaf blades are elliptic to slightly oblanceolate, and from 50 to 80 mm long (sometimes 30–100 mm long) by 20–35 mm broad (with the juvenile foliage being larger). The base of the blade is acute and attenuates to the stalk. The leaf margins have irregular and very shallowly rounded teeth, and are yellow-green or dark green above, and light green below. The margins are very minutely fringed, and there are appressed hairs on the midrib beneath. The inflorescence is a spike and is terminal on the side branches. It has a covering of long soft weak hairs which are clearly separated but not sparse and is 2–4 cm long. The inflorescence stalk is 1–2 cm long. There are four brownish red tepals (7–9 mm by 1.8–2.1 mm), and four stamens. The anther filaments are 0.7–1.5 mm long. There are 20-90 white or pale yellow flowers in an inflorescence. The perianth is a pale amber and the flowers are enclosed by shining pink, sharp-pointed bracts. The fruit is enclosed in a bladder-like sac which does not open at maturity to release the seed. It is cylindrical and 2–2.5 mm by 1–1.5 mm. It flowers and fruits from November to March.


Habitat

It appears to prefer growing by streams and in damp forests.


Taxonomy and naming

It was first described by Robert Brown in 1810, and he gave it the specific epithet, ''arborescens'', from the Latin, ''arborescens,'' "becoming a tree" or "tree-like". Conservatoire botanique national de Brest-Achyanthes arborescens-15 07 04 CP-01 (20032334208).jpg File:Conservatoire botanique national de Brest-Achyanthes arborescens-15 07 03 El Funcionario-01 (19395956941).jpg File:Conservatoire botanique national de Brest-Achyanthes arborescens-15 07 04 Philweb-05 (20051398169).jpg File:Conservatoire botanique national de Brest-Achyanthes arborescens-15 07 04 Philweb-05 (19227298828).jpg


Conservation status

''A. arborescens'' has been declared "critically endangered" under the EPBC Act. Despite setting copious seed, seedling recruitment is reduced by cattle grazing, by high levels of seed predation from the introduced rat population, and by introduced weeds such as ''
Solanum mauritianum ''Solanum mauritianum'' is a small tree or shrub native to South America, including Northern Argentina, Southern Brazil, Paraguay and Uruguay. Its common names include earleaf nightshade (or "ear-leaved nightshade"), woolly nightshade, flannel we ...
'' and ''
Homalanthus populifolius ''Homalanthus populifolius'', the bleeding heart, native poplar or Queensland poplar, is an Australian rainforest plant in the family Euphorbiaceae. It often appears in areas of rainforest disturbance. Bleeding heart is highly regarded by rainfor ...
'' which require a similar habitat to ''A. arborescens'' and possibly may out-compete it. Additionally the introduced vine ''
Ipomoea cairica ''Ipomoea cairica'' is a vining, herbaceous, perennial plant with palmate leaves and large, showy white to lavender flowers. A species of morning glory, it has many common names, including mile-a-minute vine, Messina creeper, Cairo morning glo ...
'' sometimes smothers the trees. Tourism also negatively impacts this tiny population.


References


External links

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''Achyranthes arborescens'' conservation adviceNorfolk Island Regional Recovery Plan
{{DEFAULTSORT:Achyranthes arborescens arborescens Taxa named by Robert Brown (botanist, born 1773) Plants described in 1810