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''Achlys'' is a small genus of flowering plants in the barberry family (
Berberidaceae The Berberidaceae are a family of 18 genera of flowering plants commonly called the barberry family. This family is in the order Ranunculales. The family contains about 700 known species, of which the majority are in ''Berberis''. The species i ...
), which it shares with genera such as ''
Berberis ''Berberis'' (), commonly known as barberry, is a large genus of deciduous and evergreen shrubs from tall, found throughout Temperateness, temperate and subtropical regions of the world (apart from Australia). Species diversity is greatest in S ...
'' and '' Vancouveria''.Gilkey, Helen M. & Dennis, La Rea J. (2001). ''Handbook of Northwestern Plants''. Oregon State University Press. p. 161 It is named after the Greek legendary figure associated with shade and mist,
Achlys Achlys ( grc, Ἀχλύς "mist"), in the Hesiodic ''Shield of Heracles'', is one of the figures depicted on Heracles' shield, perhaps representing the personification of sorrow. In Homer, ''achlys'' is the mist which fogs or blinds mortal eye ...
, because the plants grow in the shade.


Species

There are either two or three species, depending on the authority. ''Achlys triphylla'' and ''Achlys californica'' are both native to western North America. Another ''Achlys'' is found in Japan: some authorities treat this as a subspecies of ''A. triphylla'', while others, especially in older treatments, call this ''Achlys japonica''. Still others consider ''A. triphylla'' and ''A. californica'' too similar to be separate species. The common names for these plants include vanilla leaf (sometimes written as vanilla-leaf or vanillaleaf, depending on the taxonomist or flora), deer's foot and sweet after death, referring to the vanilla scent of its crushed leaves.Pojar & MacKinnon (2004). ''Plants of the Pacific Northwest Coast'' (Revised Edition). Lone Pine Publishing. p. 312
The Plant List The Plant List was a list of botanical names of species of plants created by the Royal Botanic Gardens, Kew and the Missouri Botanical Garden and launched in 2010. It was intended to be a comprehensive record of all known names of plant species ...
recognizes two species, with ''A. californica'' regarded as a subspecies of ''A. triphylla''


Description

''Achlys triphylla'', known in western North America as vanillaleaf, is an erect perennial plant that sprouts from a creeping
rhizome In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow hori ...
. Leaves are long-petioled and palmately divided into three leaflets. Flowers are small and lack sepals and petals, but instead have long showy white
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...
s that form single erect spikes. The leaflets give a great hint to the identification of the plant: bend back the middle leaflet and you have an upside-down set of moose antlers. Alternatively, bend back the two side leaflets and you have a goose or deer foot (hence the common name). In the Pacific Northwest, ''Achlys triphylla'' is ubiquitous in moist shady forests west of the Cascades at low to middle elevations from Vancouver island and southern
British Columbia British Columbia (commonly abbreviated as BC) is the westernmost province of Canada, situated between the Pacific Ocean and the Rocky Mountains. It has a diverse geography, with rugged landscapes that include rocky coastlines, sandy beaches, ...
south to northern
California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...
. The plants are spaced widely on the rhizomes, but often overlap in large networks that result in carpets of ''Achlys'' that dominate the near-surface understory. ''Achlys'' seems to prefer moist soil, so at middle to higher elevations it is easier to find them along streambanks or well-shaded ravines. File:Achlys japonica 5.JPG, ''Achlys japonica'' flowers File:Achlys triphylla 0973.JPG, Covered with fine hairs, ''Achlys triphylla'' achenes are concave toward stem and fleshy. File:Achlys triphylla 15379.JPG, ''Achlys triphylla'' at Standup Creek Trail 1369 Wenatchee National Forest File:Achlys japonica 6.JPG, ''Achlys japonica'' growing in Kitakami Mountains, Iwate prefacture, Japan


Insect repellent

When dried properly, the plants are strongly aromatic and smell of vanilla. Besides serving as an excellent tent
air freshener Air fresheners are consumer products that typically emit fragrance and are used in homes or commercial interiors such as restrooms, foyers, hallways, vestibules and other smaller indoor areas, as well as larger areas such as hotel lobbies, auto d ...
, ''Achlys'' has been used by native tribes of southern British Columbia as an
insect repellent An insect repellent (also commonly called "bug spray") is a substance applied to skin, clothing, or other surfaces to discourage insects (and arthropods in general) from landing or climbing on that surface. Insect repellents help prevent and cont ...
. The dried leaves were hung in bunches in doorways to ward off flies and
mosquitoe Mosquitoes (or mosquitos) are members of a group of almost 3,600 species of small flies within the family Culicidae (from the Latin ''culex'' meaning "gnat"). The word "mosquito" (formed by ''mosca'' and diminutive ''-ito'') is Spanish for "lit ...
s, and it's not unheard of for naturalists to rub the dried or even fresh leaves on exposed skin when hiking the Olympics or Cascades during the summer mosquito season. Japanese ''Achlys'' are quite similar to those found in western North America.


References


External links

*
Jepson Manual eFlora (TJM2) treatment of ''Achlys''
{{Taxonbar, from=Q4673829 Berberidaceae Berberidaceae genera Flora of British Columbia Flora of California Flora of the West Coast of the United States Insect repellents Plant toxin insecticides