''Acer dettermani'' is an extinct

maple ''Acer'' () is a genus of trees and shrubs commonly known as maples. The genus is placed in the family Sapindaceae.Stevens, P. F. (2001 onwards). Angiosperm Phylogeny Website. Version 9, June 2008 nd more or less continuously updated since http ...

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

in the family

Sapindaceae The Sapindaceae are a family of flowering plants in the order Sapindales known as the soapberry family. It contains 138 genera and 1858 accepted species. Examples include horse chestnut, maples, ackee and lychee. The Sapindaceae occur in tempera ...

described from a series of isolated

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...

. The species is known from the

late Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', "daw ...

early Oligocene The Rupelian is, in the geologic timescale, the older of two ages or the lower of two stages of the Oligocene Epoch/Series. It spans the time between . It is preceded by the Priabonian Stage (part of the Eocene) and is followed by the Chatti ...

sediments exposed in the state of

Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...

, USA. It is one of several extinct species placed in the living section '' Macrantha''.

History and classification

''Acer dettermani'' is represented by a series of fossil specimens that were recovered from late Eocene to early Oligocene outcrops of the lower Meshik Volcanics, formerly called the Meshik Formation. The rocks were first named in 1929 by R.S. Knappen for material exposed along the

Meshik River The Meshik River is a stream, long, on the Alaska Peninsula in the U.S. state of Alaska. Beginning on the flanks of Mount Aniakchak in Aniakchak National Monument and Preserve, it flows generally west into Bristol Bay near the city of Port Heide ...

near Meshik Lake. In 1996 the sequence was reexamined and subsequently renamed the Meshick Volcanics by Robert L. Detterman and a group of researchers, to better reflect the

lithology The lithology of a rock unit is a description of its physical characteristics visible at outcrop, in hand or core samples, or with low magnification microscopy. Physical characteristics include colour, texture, grain size, and composition. Lit ...

of the rocks. The age of the formation is considered to range from approximately to approximately . The rock section from which the fossils of ''Acer dettermani'' and a number of other plant fossils were recovered is a

tuff Tuff is a type of rock made of volcanic ash ejected from a vent during a volcanic eruption. Following ejection and deposition, the ash is lithified into a solid rock. Rock that contains greater than 75% ash is considered tuff, while rock cont ...

layer that is exposed on the south flank of

Mount Aniakchak Mount Aniakchak (russian: Аниакчак) is a 3,700-year-old volcanic caldera approximately in diameter, located in the Aleutian Range of Alaska, United States. Although a stratovolcano by composition, the pre-existing mountain collapsed in ...

's crater. Wolfe and Tanai interpreted the floral assemblage of Aniakchak as being a cool mesothermal forest composed of broad-leaf plants. The species was described from a pair of

type specimen In biology, a type is a particular wiktionary:en:specimen, specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to a ...

s, the

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...

leaf, specimen USNM 396014, and one

paratype In zoology and botany, a paratype is a specimen of an organism that helps define what the scientific name of a species and other taxon actually represents, but it is not the holotype (and in botany is also neither an isotype nor a syntype). Of ...

leaf, specimen USNM 396015. Both of the type specimens are currently preserved in the

paleobotanical Paleobotany, which is also spelled as palaeobotany, is the branch of botany dealing with the recovery and identification of plant remains from geological contexts, and their use for the biological reconstruction of past environments (paleogeogr ...

collections housed at the

National Museum of Natural History The National Museum of Natural History is a natural history museum administered by the Smithsonian Institution, located on the National Mall in Washington, D.C., United States. It has free admission and is open 364 days a year. In 2021, with 7 ...

, part of the

Smithsonian Institution The Smithsonian Institution ( ), or simply the Smithsonian, is a group of museums and education and research centers, the largest such complex in the world, created by the U.S. government "for the increase and diffusion of knowledge". Founded ...

Washington, D.C. ) , image_skyline = , image_caption = Clockwise from top left: the Washington Monument and Lincoln Memorial on the National Mall, United States Capitol, Logan Circle, Jefferson Memorial, White House, Adams Morgan, ...

The specimens were studied by

paleobotanist Paleobotany, which is also spelled as palaeobotany, is the branch of botany dealing with the recovery and identification of plant remains from geological contexts, and their use for the biological reconstruction of past environments (paleogeogr ...

Jack A. Wolfe Jack Albert Wolfe (1936–2005) was a United States Geological Survey paleobotanist and paleoclimatologist best known for his studies of Tertiary climate in western North America through analysis of fossil A fossil (from Classical Latin ...

of the United States Geological Survey, Denver office and Toshimasa Tanai of

Hokkaido University , or , is a Japanese national university in Sapporo, Hokkaido. It was the fifth Imperial University in Japan, which were established to be the nation's finest institutions of higher education or research. Hokkaido University is considered ...

. Wolfe and Tanai published their 1987

type description A species description is a formal description of a newly discovered species, usually in the form of a scientific paper. Its purpose is to give a clear description of a new species of organism and explain how it differs from species that have be ...

for ''A. dettermani'' in the ''Journal of the Faculty of Science, Hokkaido University''. The

etymology Etymology ()The New Oxford Dictionary of English (1998) – p. 633 "Etymology /ˌɛtɪˈmɒlədʒi/ the study of the class in words and the way their meanings have changed throughout time". is the study of the history of the Phonological chan ...

of the chosen

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

''dettermani'' is in recognition of Robert L. Detterman for his many contributions to the understanding of the

Alaska Peninsula The Alaska Peninsula (also called Aleut Peninsula or Aleutian Peninsula, ale, Alasxix̂; Sugpiaq: ''Aluuwiq'', ''Al'uwiq'') is a peninsula extending about to the southwest from the mainland of Alaska and ending in the Aleutian Islands. The ...

's Tertiary stratigraphy.

Description

Leaves of ''Acer dettermani'' are simple in structure, with perfectly actinodromous vein structure, and are generally widely elliptic in shape. The leaves are shallowly three-lobed with the lateral lobes being about one half as long as the median lobe and all lobes being triangular in outline. The leaves have three primary veins, seven to eight secondary veins, and range between an estimated long by wide in overall dimensions. The overall morphology of ''A. dettermani'' suggests placement into the ''Acer'' section ''Macrantha''. This is based on the shallow lobing with small uniformly sized teeth and the vein structure of the leaves.

References

{{Taxonbar, from=Q4673150 dettermani Plants described in 1987 Fossil taxa described in 1987 Eocene plants Oligocene plants Eocene species first appearances Rupelian species extinctions Extinct flora of North America