Acarnidae is a family of

sponge Sponges, the members of the phylum Porifera (; meaning 'pore bearer'), are a basal animal clade as a sister of the diploblasts. They are multicellular organisms that have bodies full of pores and channels allowing water to circulate through t ...

s belonging to the order

Poecilosclerida Poecilosclerida is an order of the demosponge class. It is the most speciose demosponge order with over 2200 species (World Porifera Database). It contains about 25 recognised families. They are characterised by having chelae microscleres, that i ...

. It has a global distribution, although several genera occur primarily in colder temperate waters, and several have very restricted ranges. It is estimated that there are several hundred species.

Description

Species form encrusting, massive, fan-shaped or digitate (small, un-splitting branches growing from a spreading base) growth forms that may burrow into the substrate. The species of many genera produce fistules. The outer skeleton is made of tylotes (long, slender megascleres with knobs at both ends) which may be modified to have a spined base as well as strongyles (megascleres with both ends blunt or rounded). These form tangential and/or paratangential tracts, often with an irregular or halichondroid arrangement. The inner skeleton is made of styles (megascleres with end pointed and the other end rounded) or modified styles. These form massive networks to create isodictyal (triangular meshes in which each side is one spicule long), isotropic (no differentiation into primary or secondary tracts, lines or fibers), anisotropic (primary and secondary tracts, lines or fibers) or more irregular skeletons. In encrusting forms they form plumose (skeletal structures radiate from primary fibres or spicule tracts) or hymedesmioid (monoaxon megascleres arranged singly with the heads fixed to a spongin base with the points directed outwards) skeletons instead. Spined spicules may be present or absent. When present, they take the form of acanthostyles (spiny styles) and/or unique cladotylotes (megascleres with 4-8 or even more axial branches and tubercles). Spined microscleres include palmate isochelaes and various toxas (microscleres that are curved like a bow). Both may, however, be lost in some species. Some genera also have other microscleres such as modified anisochelae such as the bipocillae (chellae with fused wings) of ''Iophon''), microrhabd-like spicules (modified microxeas or microstrongyles) and diamond-shaped microxeas (modified toxas).

Genera

The following genera are recognised: * '' Acanthorhabdus'' (Burton, 1929) * '' Acarnus'' (Gray, 1867) * '' Acheliderma'' (Topsent, 1892) *'' Cornulella'' (Dendy, 1922) *'' Cornulum'' (Carter, 1876) *'' Damiria'' (Keller, 1891) *'' Dolichacantha'' (Hentschel, 1914) *''

Iophon Iophon ( grc-gre, Ἰοφῶν, fl. 428 BC – 405 BC) was a Greek tragic poet and son of Sophocles. Iophon gained the second prize in tragic competition in 428 BC, Euripides being first, and Ion third. He must have been alive in 405 BC, the ...

'' (Gray, 1867) *'' Megaciella'' (Hallmann, 1920) *'' Paracornulum'' (Hallmann, 1920) *'' Tedaniphorbas'' (Laubenfels, 1936) *' (de Laubenfels, 1936) *'' Zyzzya'' (Laubenfels, 1936)

References

{{Taxonbar, from=Q2822638 Poecilosclerida Sponge families Demospongiae