Abies Abies
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''Abies alba'', the European silver fir or silver fir, is a
fir Firs (''Abies'') are a genus of 48–56 species of evergreen coniferous trees in the family (biology), family Pinaceae. They are found on mountains throughout much of North America, North and Central America, Europe, Asia, and North Africa. The ...
native to the
mountain A mountain is an elevated portion of the Earth's crust, generally with steep sides that show significant exposed bedrock. Although definitions vary, a mountain may differ from a plateau in having a limited Summit (topography), summit area, and ...
s of
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
, from the
Pyrenees The Pyrenees (; es, Pirineos ; french: Pyrénées ; ca, Pirineu ; eu, Pirinioak ; oc, Pirenèus ; an, Pirineus) is a mountain range straddling the border of France and Spain. It extends nearly from its union with the Cantabrian Mountains to C ...
north to
Normandy Normandy (; french: link=no, Normandie ; nrf, Normaundie, Nouormandie ; from Old French , plural of ''Normant'', originally from the word for "northman" in several Scandinavian languages) is a geographical and cultural region in Northwestern ...
, east to the
Alps The Alps () ; german: Alpen ; it, Alpi ; rm, Alps ; sl, Alpe . are the highest and most extensive mountain range system that lies entirely in Europe, stretching approximately across seven Alpine countries (from west to east): France, Sw ...
and the
Carpathians The Carpathian Mountains or Carpathians () are a range of mountains forming an arc across Central Europe. Roughly long, it is the third-longest European mountain range after the Ural Mountains, Urals at and the Scandinavian Mountains at . The ...
,
Slovenia Slovenia ( ; sl, Slovenija ), officially the Republic of Slovenia (Slovene: , abbr.: ''RS''), is a country in Central Europe. It is bordered by Italy to the west, Austria to the north, Hungary to the northeast, Croatia to the southeast, an ...
,
Croatia , image_flag = Flag of Croatia.svg , image_coat = Coat of arms of Croatia.svg , anthem = "Lijepa naša domovino"("Our Beautiful Homeland") , image_map = , map_caption = , capit ...
,
Bosnia and Herzegovina Bosnia and Herzegovina ( sh, / , ), abbreviated BiH () or B&H, sometimes called Bosnia–Herzegovina and often known informally as Bosnia, is a country at the crossroads of south and southeast Europe, located in the Balkans. Bosnia and H ...
,
Montenegro ) , image_map = Europe-Montenegro.svg , map_caption = , image_map2 = , capital = Podgorica , coordinates = , largest_city = capital , official_languages = M ...
,
Serbia Serbia (, ; Serbian language, Serbian: , , ), officially the Republic of Serbia (Serbian language, Serbian: , , ), is a landlocked country in Southeast Europe, Southeastern and Central Europe, situated at the crossroads of the Pannonian Bas ...
, and south to
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...
,
Bulgaria Bulgaria (; bg, България, Bǎlgariya), officially the Republic of Bulgaria,, ) is a country in Southeast Europe. It is situated on the eastern flank of the Balkans, and is bordered by Romania to the north, Serbia and North Macedon ...
,
Kosovo Kosovo ( sq, Kosova or ; sr-Cyrl, Косово ), officially the Republic of Kosovo ( sq, Republika e Kosovës, links=no; sr, Република Косово, Republika Kosovo, links=no), is a partially recognised state in Southeast Euro ...
,
Albania Albania ( ; sq, Shqipëri or ), or , also or . officially the Republic of Albania ( sq, Republika e Shqipërisë), is a country in Southeastern Europe. It is located on the Adriatic and Ionian Seas within the Mediterranean Sea and shares ...
and northern
Greece Greece,, or , romanized: ', officially the Hellenic Republic, is a country in Southeast Europe. It is situated on the southern tip of the Balkans, and is located at the crossroads of Europe, Asia, and Africa. Greece shares land borders with ...
; it is also commonly grown on Christmas tree plantations in the North East region of North America spanning New England in the US to the Maritime provinces of Canada.


Description

''Abies alba'' is a large
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, which ...
conifer Conifers are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single ...
ous
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
growing to tall and with a trunk diameter up to . The largest measured tree was tall and had a trunk diameter of . It occurs at altitudes of (mainly over ), on mountains with rainfall over per year. The
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
are needle-like, flattened, long and wide by thick, glossy dark green above, and with two greenish-white bands of
stomata In botany, a stoma (from Greek ''στόμα'', "mouth", plural "stomata"), also called a stomate (plural "stomates"), is a pore found in the epidermis of leaves, stems, and other organs, that controls the rate of gas exchange. The pore is bor ...
below. The leaf is usually slightly notched at the tip. The
cones A cone is a three-dimensional geometric shape that tapers smoothly from a flat base (frequently, though not necessarily, circular) to a point called the apex or vertex. A cone is formed by a set of line segments, half-lines, or lines conn ...
are long and broad, with about 150-200 scales, each scale with an exserted bract and two winged
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...
s; they disintegrate when mature to release the seeds. The wood is white, leading to the species name ''alba''. When cultivated on Christmas Tree plantations, the tree naturally forms a symmetrical triangle shape. The trees are full and dense with smell of resin, and are known to be one of the longest lasting after being cut. In the forest the evergreen tends to form stands with other firs and beeches. It is closely related to Bulgarian fir (''
Abies borisiiregis ''Abies borisii-regis'' (Bulgarian fir) is a species of fir native to the mountains of the Balkan Peninsula in Bulgaria, northern Greece, Kosovo, North Macedonia, Albania and Serbia. It occurs at altitudes of 800–1,800 m, on mounta ...
'') further to the southeast in the Balkan Peninsula,
Spanish fir ''Abies pinsapo'', the Spanish fir, is a species of tree in the family Pinaceae, native to southern Spain and northern Morocco. Related to other species of Mediterranean firs, it appears at altitudes of in the Sierra de Grazalema in the Province ...
(''
Abies pinsapo ''Abies pinsapo'', the Spanish fir, is a species of tree in the family Pinaceae, native to southern Spain and northern Morocco. Related to other species of Mediterranean firs, it appears at altitudes of in the Sierra de Grazalema in the Province ...
'') of Spain and Morocco and Sicilian fir (''
Abies nebrodensis ''Abies nebrodensis'', the Sicilian fir, is a fir native to the Madonie mountains in northern Sicily. Taxonomy It is closely related to silver fir, ''Abies alba'', which replaces it in the Apennine Mountains of Italy and elsewhere further no ...
'') in
Sicily (man) it, Siciliana (woman) , population_note = , population_blank1_title = , population_blank1 = , demographics_type1 = Ethnicity , demographics1_footnotes = , demographi ...
, differing from these and other related Euro-Mediterranean firs in the sparser foliage, with the leaves spread either side of the shoot, leaving the shoot readily visible from above. Some botanists treat Bulgarian fir and Sicilian fir as varieties of silver fir, as ''A. alba'' var. ''acutifolia'' and ''A. alba'' var. ''nebrodensis'', respectively.


Ecology

Silver fir is an important component species in the
dinaric calcareous block fir forest The dinaric calcareous silver fir forests are an endemic vegetation type of the littoral Dinaric Alps, located in the Dinaric Mountains mixed forests ecoregion in southeastern Europe. Pure stands of dinaric calcareous silver fir (''Abies alba'' ...
in the western
Balkan Peninsula The Balkans ( ), also known as the Balkan Peninsula, is a geographical area in southeastern Europe with various geographical and historical definitions. The region takes its name from the Balkan Mountains that stretch throughout the who ...
. In
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...
, the silver fir is an important component of the mixed broadleaved-coniferous forest of the Apennine Mountains, especially in northern Apennine. The fir prefer a cold and humid climate, in northern exposition, with a high rainfall (over 1500 mm per year). In the oriental Alps of Italy, silver firs grow in mixed forests with Norway spruce, beech, and other trees. Its cone scales are eaten by the
caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder Sym ...
s of the
tortrix moth The Tortricidae are a family of moths, commonly known as tortrix moths or leafroller moths, in the order Lepidoptera. This large family has over 11,000 species described, and is the sole member of the superfamily Tortricoidea, although the genus ...
''
Cydia illutana __NOTOC__ ''Cydia illutana'' is a small moth of the family Tortricidae. It is found from western and central Europe (Great Britain, the Netherlands, Austria, Germany and France), north to Scandinavia (Denmark, Norway, Sweden and Finland) and east ...
'', while '' C. duplicana'' feeds on the bark around injuries or
canker A plant canker is a small area of dead tissue, which grows slowly, often over years. Some cankers are of only minor consequence, but others are ultimately lethal and therefore can have major economic implications for agriculture and horticultur ...
.


Chemistry and pharmacology

The bark and wood of silver fir are rich in antioxidative
polyphenol Polyphenols () are a large family of naturally occurring organic compounds characterized by multiples of phenol units. They are abundant in plants and structurally diverse. Polyphenols include flavonoids, tannic acid, and ellagitannin, some of ...
s. Six phenolic acids were identified ( gallic, homovanillic, protocatechuic, p-hydroxybenzoic, vanillic and p-coumaric), three
flavonoid Flavonoids (or bioflavonoids; from the Latin word ''flavus'', meaning yellow, their color in nature) are a class of polyphenolic secondary metabolites found in plants, and thus commonly consumed in the diets of humans. Chemically, flavonoids ...
s (
catechin Catechin is a flavan-3-ol, a type of secondary metabolite providing antioxidant roles in plants. It belongs to the subgroup of polyphenols called flavonoids. The name of the catechin chemical family derives from ''catechu'', which is the tannic ...
,
epicatechin Catechin is a flavan-3-ol, a type of secondary metabolite providing antioxidant roles in plants. It belongs to the subgroup of polyphenols called flavonoids. The name of the catechin chemical family derives from ''catechu'', which is the tannic ...
and catechin tetramethyl ether) and eight
lignan The lignans are a large group of low molecular weight polyphenols found in plants, particularly seeds, whole grains, and vegetables. The name derives from the Latin word for "wood". Lignans are precursors to phytoestrogens. They may play a role ...
s ( taxiresinol, 7-(2-methyl-3,4-dihydroxytetrahydropyran-5-yloxy)-taxiresinol,
secoisolariciresinol Secoisolariciresinol is an organic compound. It is classified as a lignan, i.e., a type of phenylpropanoid. It is present in some cereals, e.g. rye, and together with matairesinol, has attracted much attention for its beneficial nutritional effect ...
,
lariciresinol Lariciresinol is a lignan, i.e., a type of phenylpropanoids. It is the precursor to enterolignans by the action of gut microflora. Enterolignans are of interest because they are speculated to exhibit beneficial medicinal properties. Occurrence I ...
,
hydroxymatairesinol Hydroxymatairesinol (HMR) is a lignan found in Norway spruce (''Picea abies''). It is an enterolactone precursor with anticancer activities. In rats, HMR decreased the volume of induced tumours and stabilised established tumours, as well as preve ...
, isolariciresinol,
matairesinol Matairesinol is an organic compound. It is classified as a lignan, i.e., a type of phenylpropanoid. It is present in some cereals, e.g. rye, and together with Secoisolariciresinol, has attracted much attention for its beneficial nutritional effec ...
and
pinoresinol Pinoresinol is a tetrahydrofuran lignan found in ''Styrax sp.'', ''Forsythia suspensa, and in Forsythia koreana''. It is also found in the caterpillar of the cabbage butterfly, ''Pieris rapae'' where it serves as a defence against ants. In food, i ...
). The extract from the trunk was shown to prevent
atherosclerosis Atherosclerosis is a pattern of the disease arteriosclerosis in which the wall of the artery develops abnormalities, called lesions. These lesions may lead to narrowing due to the buildup of atheroma, atheromatous plaque. At onset there are usu ...
in guinea pigs and to have cardioprotective effect in isolated rat hearts. Silver fir wood extract was found to reduce the post-prandial glycemic response (concentration of sugar in the blood after the meal) in healthy volunteers.


Uses

In
Roman times In modern historiography, ancient Rome refers to Roman civilisation from the founding of the city of Rome in the 8th century BC to the collapse of the Western Roman Empire in the 5th century AD. It encompasses the Roman Kingdom (753–509 BC ...
the wood was used to make wooden casks to store and transport wine and other substances. A resinous
essential oil An essential oil is a concentrated hydrophobic liquid containing volatile (easily evaporated at normal temperatures) chemical compounds from plants. Essential oils are also known as volatile oils, ethereal oils, aetheroleum, or simply as the o ...
can be extracted. This pine-scented oil is used in perfumes, bath products, and aerosol inhalants. Its branches (including the leaves, bark and wood) were used for production of
spruce beer Spruce beer is a beverage flavored with the buds, needles, or essence of spruce trees. ''Spruce beer'' can refer to either alcoholic or non-alcoholic beverages. A number of flavors are associated with spruce-flavored beverages, ranging from flo ...
. Silver fir is the species first used as a
Christmas tree A Christmas tree is a decorated tree, usually an evergreen conifer, such as a spruce, pine or fir, or an artificial tree of similar appearance, associated with the celebration of Christmas. The custom was further developed in early modern ...
, but has been largely replaced by
Nordmann fir ''Abies nordmanniana'', the Nordmann fir or Caucasian fir, is a fir Indigenous (ecology), indigenous to the mountains south and east of the Black Sea, in Turkey, Georgia (country), Georgia and the Russian Caucasus. It occurs at altitudes of 900 ...
(which has denser, more attractive foliage),
Norway spruce ''Picea abies'', the Norway spruce or European spruce, is a species of spruce native to Northern, Central and Eastern Europe. It has branchlets that typically hang downwards, and the largest cones of any spruce, 9–17 cm long. It is very close ...
(which is much cheaper to grow), and other species. The
wood Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of lignin th ...
is strong, lightweight, light-colored, fine grained, even-textured and long fibered. The
timber Lumber is wood that has been processed into dimensional lumber, including beams and planks or boards, a stage in the process of wood production. Lumber is mainly used for construction framing, as well as finishing (floors, wall panels, wi ...
is mainly used as construction wood, furniture,
plywood Plywood is a material manufactured from thin layers or "plies" of wood veneer that are glued together with adjacent layers having their wood grain rotated up to 90 degrees to one another. It is an engineered wood from the family of manufactured ...
,
pulpwood Pulpwood is timber with the principal use of making wood pulp for paper production. Applications * Trees raised specifically for pulp production account for 15% of world pulp production, old growth forests 9% and second- and third- and more gener ...
and
paper Paper is a thin sheet material produced by mechanically or chemically processing cellulose fibres derived from wood, rags, grasses or other vegetable sources in water, draining the water through fine mesh leaving the fibre evenly distributed ...
manufacture. The honeydew which is produced by
aphids Aphids are small sap-sucking insects and members of the superfamily Aphidoidea. Common names include greenfly and blackfly, although individuals within a species can vary widely in color. The group includes the fluffy white woolly aphids. A t ...
sitting on the silver fir is collected by
honey bee A honey bee (also spelled honeybee) is a eusocial flying insect within the genus ''Apis'' of the bee clade, all native to Afro-Eurasia. After bees spread naturally throughout Africa and Eurasia, humans became responsible for the current co ...
s. The resulting
honey Honey is a sweet and viscous substance made by several bees, the best-known of which are honey bees. Honey is made and stored to nourish bee colonies. Bees produce honey by gathering and then refining the sugary secretions of plants (primar ...
is marketed as "fir honey".


Etymology

''Abies'' is derived from
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
, meaning 'rising one'. The name was used to refer to tall trees or ships.Gledhill, David (2008). "The Names of Plants". Cambridge University Press. (hardback), (paperback). pp 32, 41 ''Alba'' means 'bright' or 'dead white'.


References

*


External links


conifers.org: ''Abies alba''botany.cz: ''Abies alba'' Millphotomazza.com: ''Abies alba''conifersaroundtheworld.com: Abies alba - European White Fir

pfaf.org: ''Abies alba'' Mill.monumentaltrees.com: The thickest, tallest, and oldest European silver fir trees (''Abies alba'')baumkunde.de: Weiß-Tanne (''Abies alba'')
, In German
''Abies alba''
Distribution map, genetic conservation units and related resources.
European Forest Genetic Resources Programme European Forest Genetic Resources Programme (EUFORGEN) is an international network that supports the conservation and sustainable use of forest genetic resources in Europe. The programme’s tasks include to coordinate and promote ''in situ ' ...
(EUFORGEN) {{Authority control
alba ''Alba'' ( , ) is the Scottish Gaelic name for Scotland. It is also, in English language historiography, used to refer to the polity of Picts and Scottish people, Scots united in the ninth century as the Kingdom of Alba, until it developed i ...
Trees of Europe Flora of the Alps Flora of Spain Flora of the Pyrenees Flora of France Flora of the Carpathians Plants described in 1768 Flora of Montenegro Taxa named by Philip Miller