The alula , or bastard wing, (plural ''alulae'') is a small projection on the

anterior Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position prov ...

edge of the

wing A wing is a type of fin that produces lift while moving through air or some other fluid. Accordingly, wings have streamlined cross-sections that are subject to aerodynamic forces and act as airfoils. A wing's aerodynamic efficiency is expres ...

of modern

bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...

s and a few non-avian dinosaurs. The word is

Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...

and means "winglet"; it is the

diminutive A diminutive is a root word that has been modified to convey a slighter degree of its root meaning, either to convey the smallness of the object or quality named, or to convey a sense of intimacy or endearment. A (abbreviated ) is a word-formati ...

of ''ala'', meaning "wing". The alula is the freely moving first digit, a bird's "thumb", and typically bears three to five small

flight feather Flight feathers (''Pennae volatus'') are the long, stiff, asymmetrically shaped, but symmetrically paired pennaceous feathers on the wings or tail of a bird; those on the wings are called remiges (), singular remex (), while those on the tail ...

s, with the exact number depending on the

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

. There also are minor

covert feather A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which, as the name implies, cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are sm ...

s overlying the flight feathers. Like the larger

s found on the wing's trailing edge, these alula feathers are asymmetrical, with the shaft running closer to anterior edge.

Function

In most situations, the alula is held flush against the wing; however, it can be manipulated. When flying at slow speeds or landing, the bird moves its alula slightly upwards and forward, which creates a small slot on the wing's leading edge. This functions in the same way as the

slats Super Low Altitude Test Satellite (SLATS) or Tsubame was a JAXA satellite intended to demonstrate operations in very low Earth orbit (VLEO, below 200 km), using ion engines to counteract aerodynamic drag from the Earth's atmosphere which i ...

on the wing of an aircraft, allowing the wing to achieve a higher than normal

angle of attack In fluid dynamics, angle of attack (AOA, α, or \alpha) is the angle between a reference line on a body (often the chord line of an airfoil) and the vector representing the relative motion between the body and the fluid through which it is m ...

– and thus lift – without resulting in a stall. The tip of the alula forms a tiny vortex, acting similar to a vortex generator, that forces the airflow over the wing to better bind to it. During stretching of the wing down toward the ground, the alula is abducted from the wing and can be clearly viewed. In falcons, the alula is more prominent, and provides a degree of control when the wing is cupped or tucked in a dive/stoop. The Alulae are particularly notable in

peregrine falcon The peregrine falcon (''Falco peregrinus''), also known as the peregrine, and historically as the duck hawk in North America, is a Cosmopolitan distribution, cosmopolitan bird of prey (Bird of prey, raptor) in the family (biology), family Falco ...

In ancient birds

The presence of an alula has been confirmed in several now-

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

ancient relatives of modern birds, including '' Eoalulavis hoyasi'' (an

enantiornithine The Enantiornithes, also known as enantiornithines or enantiornitheans in literature, are a group of extinct avialans ("birds" in the broad sense), the most abundant and diverse group known from the Mesozoic era. Almost all retained teeth and cla ...

from the mid-

Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...

, 115

mya Mya may refer to: Brands and product names * Mya (program), an intelligent personal assistant created by Motorola * Mya (TV channel), an Italian Television channel * Midwest Young Artists, a comprehensive youth music program Codes * Burmese ...

) and the earlier '' Protopteryx fengningensis''. Since these

are not closely related to modern birds, either the alula

evolved Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation t ...

twice, or it did so more than 130 million years ago.

Citations

References

* * * {{Birds Feathers Bird anatomy

Function

In ancient birds

See also

Citations

References