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2019 in paleoentomology is a list of new
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (head, thorax and abdomen), three pairs ...
taxa In biology, a taxon ( back-formation from '' taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular n ...
that were described during the year 2019, as well as other significant discoveries and events related to paleoentomology that were scheduled to occur during the year.


New taxa


Coleopterans


Dictyopterans


Dipterans


Hemipterans


Hymenopterans


Mecopterans


Neuropterans


Odonatans


Orthopterans


Phasmatodea


Plecopterans


Strepsipterans


Thysanoptera


Trichopterans


Other insects


General research

* A study aiming to estimate the taxonomic diversity of insects in deep time is published by Schachat ''et al.'' (2019). * A study on the phylogenetic relationships of the Carboniferous insect '' Stephanastus polinae'' is published by Beutel, Yan & Kukalová-Peck (2019). * A study on the age and depositional environment of the Xiaheyan insect fauna (
Ningxia Ningxia (,; , ; alternately romanized as Ninghsia), officially the Ningxia Hui Autonomous Region (NHAR), is an autonomous region in the northwest of the People's Republic of China. Formerly a province, Ningxia was incorporated into Gansu in ...
, China) is published online by Trümper ''et al.'' (2019). * A study on 240-million-year-old insect fossils from the Mount San Giorgio
Lagerstätte A Lagerstätte (, from ''Lager'' 'storage, lair' '' Stätte'' 'place'; plural ''Lagerstätten'') is a sedimentary deposit that exhibits extraordinary fossils with exceptional preservation—sometimes including preserved soft tissues. These for ...
( Switzerland
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...
), evaluating their implications for the knowledge of the time scale of insect evolution, is published by Montagna ''et al.'' (2019). * A study on insect fossils from the
Upper Triassic The Late Triassic is the third and final epoch of the Triassic Period in the geologic time scale, spanning the time between Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch and followed by the Early Jurassic Epoch. T ...
Laohugou Formation (
Hebei Hebei or , (; alternately Hopeh) is a northern province of China. Hebei is China's sixth most populous province, with over 75 million people. Shijiazhuang is the capital city. The province is 96% Han Chinese, 3% Manchu, 0.8% Hui, and ...
, China) is published by Huang ''et al.'' (2019). * The earliest evidence of exophytic oviposition (eggs laid directly on the outer surface of plants) known so far, in the form of insect eggs (probably produced by roachoids) preserved on plant fossils, is reported from the Carboniferous (
Gzhelian The Gzhelian ( ) is an age in the ICS geologic timescale or a stage in the stratigraphic column. It is the youngest stage of the Pennsylvanian, the youngest subsystem of the Carboniferous. The Gzhelian lasted from to Ma. It follows the Ka ...
) of the Saale Basin (
Germany Germany, officially the Federal Republic of Germany (FRG),, is a country in Central Europe. It is the most populous member state of the European Union. Germany lies between the Baltic and North Sea to the north and the Alps to the sou ...
) by Laaß & Hauschke (2019). * A study on the frequency and diversity of damage types caused by insect oviposition in plants from the
Upper Triassic The Late Triassic is the third and final epoch of the Triassic Period in the geologic time scale, spanning the time between Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch and followed by the Early Jurassic Epoch. T ...
Yangcaogou Formation,
Middle Jurassic The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations ...
Jiulongshan Formation The Haifanggou Formation (), also known as the Jiulongshan Formation (), is a fossil-bearing rock deposit located near Daohugou () village of Ningcheng County, in Inner Mongolia, northeastern China. The formation consists of coarse Conglomerate (g ...
and
Lower Cretaceous Lower may refer to: *Lower (surname) *Lower Township, New Jersey *Lower Receiver (firearms) *Lower Wick Gloucestershire, England See also *Nizhny Nizhny (russian: Ни́жний; masculine), Nizhnyaya (; feminine), or Nizhneye (russian: Ни́� ...
Yixian Formation The Yixian Formation (; formerly transcribed as Yihsien Formation) is a geological formation in Jinzhou, Liaoning, People's Republic of China, that spans the late Barremian and early Aptian stages of the Early Cretaceous. It is known for its ...
( China), assessing the degree of plant host specificity, is published by Lin ''et al.'' (2019). * A study on the plant specimens (ferns, gymnosperms and angiosperms) from the
Lower Cretaceous Lower may refer to: *Lower (surname) *Lower Township, New Jersey *Lower Receiver (firearms) *Lower Wick Gloucestershire, England See also *Nizhny Nizhny (russian: Ни́жний; masculine), Nizhnyaya (; feminine), or Nizhneye (russian: Ни́� ...
Araripe Basin The Araripe Basin () is a rift basin covering about ,Neto et al., 2013, p.1 in Ceará, Piauí and Pernambuco states of northeastern Brazil. It is bounded by the Patos and Pernambuco lineaments, and is situated east of the Parnaíba Basin, southw ...
(
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
) preserving evidence of plant–insect interactions and potentially of paleoecological relationships between plants and insects is published by Edilson Bezerra dos Santos Filho ''et al.'' (2019). * Leaves of members of the family
Nymphaeaceae Nymphaeaceae () is a family of flowering plants, commonly called water lilies. They live as rhizomatous aquatic herbs in temperate and tropical climates around the world. The family contains nine genera with about 70 known species. Water ...
preserving evidence of insect herbivory are reported from the
Albian The Albian is both an age of the geologic timescale and a stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early/Lower Cretaceous Epoch/ Series. Its approximate time range is 113.0 ± 1.0 Ma to 100.5 ± ...
Utrillas Formation Utrillas is a municipality located in the province of Teruel, Aragon Aragon ( , ; Spanish and an, Aragón ; ca, Aragó ) is an autonomous community in Spain, coextensive with the medieval Kingdom of Aragon. In northeastern Spain, the Ar ...
(
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = '' Plus ultra'' ( Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , ...
) by Estévez-Gallardo ''et al.'' (2019). * A study on changes in insect and plant communities across the
Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch that lasted from about 66 to 56 million years ago (mya). It is the first epoch of the Paleogene Period in the modern Cenozoic Era. The name is a combination of the Ancient Greek ''pal ...
Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', " ...
boundary within the Hanna Basin (
Wyoming Wyoming () is a state in the Mountain West subregion of the Western United States. It is bordered by Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho to the west, Utah to the southwest, and Colorado to the sou ...
,
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., federal district, five ma ...
) is published by Azevedo Schmidt ''et al.'' (2019). * A study on insect
pupa A pupa ( la, pupa, "doll"; plural: ''pupae'') is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in their ...
tion structures from the
Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campani ...
dinosaur nesting site at the Egg Mountain locality (
Two Medicine Formation The Two Medicine Formation is a geological formation, or rock body, in northwestern Montana and southern Alberta that was deposited between and (million years ago), during Campanian (Late Cretaceous) time. It crops out to the east of the Rocky M ...
;
Montana Montana () is a U.S. state, state in the Mountain states, Mountain West List of regions of the United States#Census Bureau-designated regions and divisions, division of the Western United States. It is bordered by Idaho to the west, North ...
,
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., federal district, five ma ...
) is published by Freimuth & Varricchio (2019). * Redescription and a study on the phylogenetic relationships of a
Permian The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Pale ...
orthoptera Orthoptera () is an order of insects that comprises the grasshoppers, locusts, and crickets, including closely related insects, such as the bush crickets or katydids and wētā. The order is subdivided into two suborders: Caelifera – grass ...
n '' Vologdoptera maculata'' is published by Aristov & Gorochov (2019). * A study on the phylogenetic relationships of the
Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch that lasted from about 66 to 56 million years ago (mya). It is the first epoch of the Paleogene Period in the modern Cenozoic Era. The name is a combination of the Ancient Greek ''pal ...
orthopteran '' Hylophalangopsis chinensis'' is published by Wang ''et al.'' (2019). * 308-million-years-old female palaeodictyopteran
nymphs A nymph ( grc, νύμφη, nýmphē, el, script=Latn, nímfi, label=Modern Greek; , ) in ancient Greek folklore is a minor female nature deity. Different from Greek goddesses, nymphs are generally regarded as personifications of nature, are ty ...
, preserving large pointing structures interpreted as
ovipositor The ovipositor is a tube-like organ used by some animals, especially insects, for the laying of eggs. In insects, an ovipositor consists of a maximum of three pairs of appendages. The details and morphology of the ovipositor vary, but typica ...
s, are described from the Piesberg quarry in northwestern
Germany Germany, officially the Federal Republic of Germany (FRG),, is a country in Central Europe. It is the most populous member state of the European Union. Germany lies between the Baltic and North Sea to the north and the Alps to the sou ...
by Kiesmüller ''et al.'' (2019). * A study on the anatomy of immature stages and adult specimens of members of Palaeodictyopterida, evaluating its implications for different hypotheses about lifestyle strategies of these insects, is published by Prokop ''et al.'' (2019). * A revision of phyloblattid and compsoblattid
blattodea Blattodea is an order of insects that contains cockroaches and termites. Formerly, termites were considered a separate order, Isoptera, but genetic and molecular evidence suggests they evolved from within the cockroach lineage, cladistically m ...
ns from the Carboniferous (
Kasimovian The Kasimovian is a geochronologic age or chronostratigraphic stage in the ICS geologic timescale. It is the third stage in the Pennsylvanian (late Carboniferous), lasting from to Ma.; 2004: ''A Geologic Time Scale 2004'', Cambridge Universi ...
) of the Souss basin (southwestern
High Atlas High Atlas, also called the Grand Atlas ( ar, الأطلس الكبير, Al-Aṭlas al-Kabīr; french: Haut Atlas; shi, ⴰⴷⵔⴰⵔ ⵏ ⴷⵔⵏ ''Adrar n Dern''), is a mountain range in central Morocco, North Africa, the highest part of t ...
mountains,
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria ...
) is published by Belahmira ''et al.'' (2019). * A fossil
ootheca An ootheca (pl. ''oothecae'' ) is a type of egg mass made by any member of a variety of species including mollusks (such as '' Turbinella laevigata''), mantises, and cockroaches. The word is a Latinized combination of ''oo-'', meaning "egg", ...
resembling those of extant
mantis Mantises are an order (Mantodea) of insects that contains over 2,400 species in about 460 genera in 33 families. The largest family is the Mantidae ("mantids"). Mantises are distributed worldwide in temperate and tropical habitats. They ha ...
es is described from the
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...
amber from
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John Wells explai ...
by Li & Huang (2019). * An assemblage of 14
dictyoptera Dictyoptera (from Greek δίκτυον ''diktyon'' "net" and πτερόν ''pteron'' "wing") is an insect superorder that includes two extant orders of polyneopterous insects: the order Blattodea (termites and cockroaches together) and the ord ...
ns is reported from the
Santonian The Santonian is an age in the geologic timescale or a chronostratigraphic stage. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series. It spans the time between 86.3 ± 0.7 mya (million years ago) and 83.6 ± 0.7 mya. ...
amber from Yantardakh (
Taymyr Peninsula The Taymyr Peninsula (russian: Таймырский полуостров, Taymyrsky poluostrov) is a peninsula in the Far North of Russia, in the Siberian Federal District, that forms the northernmost part of the mainland of Eurasia. Administrat ...
,
Russia Russia (, , ), or the Russian Federation, is a transcontinental country spanning Eastern Europe and Northern Asia. It is the largest country in the world, with its internationally recognised territory covering , and encompassing one-eigh ...
) by Vršanský (2019). * A study on the anatomy and life habits of alienopterans is published by Wipfler ''et al.'' (2019). * A redescription of a female and a description of a male of the parvaverrucosid species ''
Parvaverrucosa ''Parvaverrucosa'' is an insect genus in the extinct, monotypic family Parvaverrucosidae, of the order Hemiptera. It contains the monotypic species ''Parvaverrucosa annulata'' known from the Cenomanian aged Burmese amber of Myanmar. First desc ...
annulata'' from the Cretaceous amber from Myanmar is published by Węgierek, Cai & Huang (2019). * A gregarious assemblage of protopsyllidioids belonging to the genus '' Postopsyllidium'' is reported from a single piece of the Cretaceous amber from Myanmar by Hakim, Azar & Huang (2019). * A redescription of the fossil
water strider The Gerridae are a family of insects in the order Hemiptera, commonly known as water striders, water skeeters, water scooters, water bugs, pond skaters, water skippers, or water skimmers. Consistent with the classification of the Gerridae as t ...
species '' Aquarius lunpolaensis'' based on new specimens from the Lunpola and Nima basins of central
Tibet Tibet (; ''Böd''; ) is a region in East Asia, covering much of the Tibetan Plateau and spanning about . It is the traditional homeland of the Tibetan people. Also resident on the plateau are some other ethnic groups such as Monpa people, ...
is published by Cai ''et al.'' (2019). * New specimens of '' Mesodiphthera grandis'' are described from the
Norian The Norian is a division of the Triassic Period. It has the rank of an age ( geochronology) or stage (chronostratigraphy). It lasted from ~227 to million years ago. It was preceded by the Carnian and succeeded by the Rhaetian. Stratigraphic ...
insect locality at Dinmore (
Queensland ) , nickname = Sunshine State , image_map = Queensland in Australia.svg , map_caption = Location of Queensland in Australia , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , established_ ...
, Australia) by Lambkin (2019), who interprets this taxon as a hairy cicada and the oldest known
cicada The cicadas () are a superfamily, the Cicadoidea, of insects in the order Hemiptera (true bugs). They are in the suborder Auchenorrhyncha, along with smaller jumping bugs such as leafhoppers and froghoppers. The superfamily is divided into two ...
reported so far. * Revision of members of extinct
cicadomorpha Cicadomorpha is an infraorder of the insect order Hemiptera which contains the cicadas, leafhoppers, treehoppers, and spittlebugs. There are approximately 35,000 described species worldwide. Distributed worldwide, all members of this group ar ...
n family Hylicellidae from the Norian
Mount Crosby Formation Mount is often used as part of the name of specific mountains, e.g. Mount Everest. Mount or Mounts may also refer to: Places * Mount, Cornwall, a village in Warleggan parish, England * Mount, Perranzabuloe, a hamlet in Perranzabuloe parish ...
(Queensland, Australia) is published by Lambkin (2019). * A group of heteropteran
nymphs A nymph ( grc, νύμφη, nýmphē, el, script=Latn, nímfi, label=Modern Greek; , ) in ancient Greek folklore is a minor female nature deity. Different from Greek goddesses, nymphs are generally regarded as personifications of nature, are ty ...
preserved together with their eggs, including two nymphs caught in the act of hatching, is described from the
Dominican amber Dominican amber is amber from the Dominican Republic derived from resin of the extinct tree ''Hymenaea protera''. Dominican amber differentiates itself from Baltic amber by being nearly always transparent, and it has a higher number of fossil incl ...
by Hörnig, Fischer & Haug (2019). * A female specimen of the myopsocid species '' Myopsocus arthuri'' is described from the
Dominican amber Dominican amber is amber from the Dominican Republic derived from resin of the extinct tree ''Hymenaea protera''. Dominican amber differentiates itself from Baltic amber by being nearly always transparent, and it has a higher number of fossil incl ...
by Hakim ''et al.'' (2019). * A study on the phylogenetic relationships of fossil
ichneumonid The Ichneumonidae, also known as the ichneumon wasps, Darwin wasps, or ichneumonids, are a family of parasitoid wasps of the insect order Hymenoptera. They are one of the most diverse groups within the Hymenoptera with roughly 25,000 species cur ...
wasps is published by Klopfstein & Spasojevic (2019), who transfer the species ''" Plectiscidea" lanhami'' to the genus ''
Allomacrus ''Allomacrus'' is a genus of parasitoid wasps belonging to the family Ichneumonidae. The species of this genus are found in Europe and Northern America. Species: * ''Allomacrus arcticus'' (Holmgren, 1880) * ''Allomacrus jakuticus'' Humala, 200 ...
''. * A fossilized hind wing of a member of the family Ichneumonidae is described from a Calabrian sandstone from
Madeira ) , anthem = ( en, "Anthem of the Autonomous Region of Madeira") , song_type = Regional anthem , image_map=EU-Portugal_with_Madeira_circled.svg , map_alt=Location of Madeira , map_caption=Location of Madeira , subdivision_type=Sovereign st ...
by Góis-Marques ''et al.'' (2019). * Revision of fossil figitids from the Late Eocene of Florissant (
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., federal district, five ma ...
) and from the Oligocene–Miocene boundary of Rott-am-Siebengebirge (
Germany Germany, officially the Federal Republic of Germany (FRG),, is a country in Central Europe. It is the most populous member state of the European Union. Germany lies between the Baltic and North Sea to the north and the Alps to the sou ...
) is published by Pujade-Villar & Peñalver (2019). * A study on the evolutionary history of
colletid The Colletidae are a family of bees, and are often referred to collectively as plasterer bees or polyester bees, due to the method of smoothing the walls of their nest cells with secretions applied with their mouthparts; these secretions dry int ...
bees belonging to the group Neopasiphaeinae, as indicated by phylogenetic,
biogeographic Biogeography is the study of the distribution of species and ecosystems in geographic space and through geological time. Organisms and biological communities often vary in a regular fashion along geographic gradients of latitude, elevation, ...
and paleontological data, is published by Almeida ''et al.'' (2019). * The first corydalid
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. Th ...
preserved with gut contents is described from the Lower Cretaceous
Yixian Formation The Yixian Formation (; formerly transcribed as Yihsien Formation) is a geological formation in Jinzhou, Liaoning, People's Republic of China, that spans the late Barremian and early Aptian stages of the Early Cretaceous. It is known for its ...
( China) by Zhao ''et al.'' (2019). * Seed of the ginkgoalean ''
Yimaia ''Yimaia'' is a extinct genus of Ginkgoalean tree, and the only member of the family Yimaiaceae. In botanical form classification, its a form taxon for ginkgoalean ovulate organs. ''Yimaia'' species are distinguished from other Ginkgoales by th ...
capituliformis'' with damage interpreted as likely oviposition lesions inflicted by a
kalligrammatid Kalligrammatidae, sometimes known as kalligrammatids or kalligrammatid lacewings, is a family of extinct insects in the order Neuroptera (lacewings) that contains twenty genera and a number of species. The family lived from the Middle Jurassic ...
lacewing The insect order (biology), order Neuroptera, or net-winged insects, includes the lacewings, Mantispidae, mantidflies, antlions, and their relatives. The order consists of some 6,000 species. Neuroptera can be grouped together with the Megalopt ...
is described from the
Middle Jurassic The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations ...
Jiulongshan Formation The Haifanggou Formation (), also known as the Jiulongshan Formation (), is a fossil-bearing rock deposit located near Daohugou () village of Ningcheng County, in Inner Mongolia, northeastern China. The formation consists of coarse Conglomerate (g ...
( China) by Meng ''et al.'' (2019). * A long-necked neuropteran larva, preserving a unique combination of anatomical characters present in various neuropterans families, is described from the Cretaceous amber from
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John Wells explai ...
by Haug ''et al.'' (2019). * A larval lacewing with unusually large mandibulo-maxillary piercing stylets is described from the Cretaceous amber from Myanmar by Haug, Müller & Haug (2019). * A neuropteran larva with prominent curved stylets is described from the Cretaceous amber from Myanmar by Haug, Müller & Haug (2019). * A late
instar An instar (, from the Latin '' īnstar'', "form", "likeness") is a developmental stage of arthropods, such as insects, between each moult (''ecdysis''), until sexual maturity is reached. Arthropods must shed the exoskeleton in order to grow o ...
strepsiptera The Strepsiptera are an order of insects with eleven extant families that include about 600 described species. They are endoparasites in other insects, such as bees, wasps, leafhoppers, silverfish, and cockroaches. Females of most species never ...
n larva, probably belonging to the genus '' Mengea'', is described from the
Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', " ...
Baltic amber The Baltic region is home to the largest known deposit of amber, called Baltic amber or succinite. It was produced sometime during the Eocene epoch, but exactly when is controversial. It has been estimated that these forests created more than 1 ...
by Pohl ''et al.'' (2019). * A review of plants, fungi and animals found associated with fossil beetles from Myanmar, Dominican and
Mexican amber Mexican amber, also known as Chiapas Amber is amber found in Mexico, created during the Early Miocene and middle Miocene epochs of the Cenozoic Era in southwestern North America. As with other ambers, a wide variety of taxa have been found as in ...
is published by Poinar (2019). * A study on the phylogenetic relationships of '' Tunguskagyrus'' is published by Beutel, Yan & Lawrence (2019). * A study on the phylogenetic relationships of '' Leehermania prorova'' is published online by Fikáček ''et al.'' (2019). * Extremely miniaturized insects interpreted as larvae of beetles belonging to the family
Ripiphoridae Ripiphoridae (formerly spelled Rhipiphoridae) is a cosmopolitan family of some 450 described species of beetles sometimes called "wedge-shaped beetles". Ripiphoridae are unusual among beetle families in that many species are hypermetamorphic par ...
and the subfamily Ripidiinae are described from the Cretaceous amber from Myanmar by Batelka ''et al.'' (2019). * A male specimen of the monotomid beetle species '' Cretakarenni birmanicus'' is described from the Cretaceous amber from Myanmar by Jiang, Liu & Wang (2019). * A protrusible prey-capture apparatus is reported in two stenine
rove beetle The rove beetles are a family (Staphylinidae) of beetles, primarily distinguished by their short elytra (wing covers) that typically leave more than half of their abdominal segments exposed. With roughly 63,000 species in thousands of genera, th ...
s from the
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...
amber from
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John Wells explai ...
(a specimen of '' Festenus gracilis'' and a specimen representing a new species of ''Festenus'') by Cai ''et al.'' (2019). * The first case of aggregation behaviour of the rove beetle '' Clidicostigus arachnipes'' is reported from the Cretaceous amber from Myanmar by Yin & Zhuo (2019). * Six well-preserved specimens of the silvanid beetle '' Protoliota'', confirming the presence of remarkable
sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most an ...
in this genus, are reported from the Cretaceous amber from Myanmar by Cai & Huang (2019). * A study on the phylogenetic relationships of extant and fossil members of the
scarab beetle The family Scarabaeidae, as currently defined, consists of over 30,000 species of beetles worldwide; they are often called scarabs or scarab beetles. The classification of this family has undergone significant change in recent years. Several sub ...
subfamily
Aclopinae Aclopinae is a subfamily of Scarabaeidae or scarab beetles in the superfamily Scarabaeoidea Scarabaeoidea is a superfamily of beetles, the only subgroup of the infraorder Scarabaeiformia. Around 35,000 species are placed in this superfamily a ...
is published by Neita‐Moreno ''et al.'' (2019). * A
flower chafer Flower chafers are a group of scarab beetles, comprising the subfamily Cetoniinae. Many species are diurnal and visit flowers for pollen and nectar, or to browse on the petals. Some species also feed on fruit. The group is also called fruit an ...
belonging to the tribe
Trichiini The Trichiini are a tribe of the scarab beetle family (Scarabaeidae), though historically they were often classified as a subfamily, Trichiinae. The conspicuous bee beetles (''Trichius'') are probably the best-known genus in Europe. They vary ...
is described from the
Baltic amber The Baltic region is home to the largest known deposit of amber, called Baltic amber or succinite. It was produced sometime during the Eocene epoch, but exactly when is controversial. It has been estimated that these forests created more than 1 ...
by Alekseev (2019), representing the earliest fossil record of the tribe Trichiini and the first known flower chafer in Baltic amber. * Two beetle larvae with unusually large terminal end compared to that in extant forms, identified as representatives of Scraptiidae, are described from the
Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', " ...
Baltic amber The Baltic region is home to the largest known deposit of amber, called Baltic amber or succinite. It was produced sometime during the Eocene epoch, but exactly when is controversial. It has been estimated that these forests created more than 1 ...
by Haug & Haug (2019). * A study comparing extant beetle fauna from
Maungatautari Maungatautari is a mountain, rural community, and ecological area near Cambridge in the Waikato region in New Zealand's central North Island. The New Zealand Ministry for Culture and Heritage gives a translation of "mountain of the upright ...
(
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 List of islands of New Zealand, smaller islands. It is the ...
) with the late
Holocene The Holocene ( ) is the current geological epoch. It began approximately 11,650 cal years Before Present (), after the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene togeth ...
beetle assemblage from two central
North Island The North Island, also officially named Te Ika-a-Māui, is one of the two main islands of New Zealand, separated from the larger but much less populous South Island by the Cook Strait. The island's area is , making it the world's 14th-larges ...
fossil sites is published by
Watts Watts is plural for ''watt'', the unit of power. Watts may also refer to: People * Watts (surname), list of people with the surname Watts Fictional characters *Watts, main character in the film '' Some Kind of Wonderful'' * Watts family, six cha ...
''et al.'' (2019). * Poisonous
setae In biology, setae (singular seta ; from the Latin word for " bristle") are any of a number of different bristle- or hair-like structures on living organisms. Animal setae Protostomes Annelid setae are stiff bristles present on the body. ...
are identified in a small caterpillar from the Eocene
Baltic amber The Baltic region is home to the largest known deposit of amber, called Baltic amber or succinite. It was produced sometime during the Eocene epoch, but exactly when is controversial. It has been estimated that these forests created more than 1 ...
by Poinar & Vega (2019). * A revision of putative fossil members of the family
Hepialidae The Hepialidae are a family of insects in the lepidopteran order. Moths of this family are often referred to as swift moths or ghost moths. Taxonomy and systematics The Hepialidae constitute by far the most diverse group of the infraorder Exo ...
is published by Simonsen, Wagner & Heikkilä (2019). * A study on the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...
specimen of the
Late Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the ...
acrocerid fly '' Archocyrtus kovalevi'' is published by Khramov & Lukashevich (2019), who report evidence of an extremely long
proboscis A proboscis () is an elongated appendage from the head of an animal, either a vertebrate or an invertebrate. In invertebrates, the term usually refers to tubular mouthparts used for feeding and sucking. In vertebrates, a proboscis is an elonga ...
, almost twice the length of the body of this insect. * A study on the phylogenetic relationships of extant and fossil acrocerid flies is published by Gillung & Winterton (2019). * A study on the phylogenetic relationships of the
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...
members of the family
Ceratopogonidae Ceratopogonidae is a family of flies commonly known as no-see-ums, or biting midges, generally in length. The family includes more than 5,000 species, distributed worldwide, apart from the Antarctic and the Arctic. Ceratopogonidae are holomet ...
is published by Borkent (2019). * A study on the morphology and phylogenetic relationships of '' Lebanoculicoides daheri'', as indicated by data from a male specimen from the Cretaceous
Lebanese Lebanese may refer to: * Something of, from, or related to the Lebanese Republic * Lebanese people The Lebanese people ( ar, الشعب اللبناني / ALA-LC: ', ) are the people inhabiting or originating from Lebanon. The term may al ...
amber, is published by Borkent (2019). * The first record of
chironomid The Chironomidae (informally known as chironomids, nonbiting midges, or lake flies) comprise a family of nematoceran flies with a global distribution. They are closely related to the Ceratopogonidae, Simuliidae, and Thaumaleidae. Many species ...
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. Th ...
from amber, comparable to larvae of modern representatives of the genus '' Bryophaenocladius'', is reported from the
Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', " ...
Baltic amber The Baltic region is home to the largest known deposit of amber, called Baltic amber or succinite. It was produced sometime during the Eocene epoch, but exactly when is controversial. It has been estimated that these forests created more than 1 ...
by Baranov ''et al.'' (2019).


References

{{Reflist, 30em 2019 in paleontology Paleoentomology