Whatcheeriid
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Whatcheeriid
Whatcheeriidae is an extinct family of tetrapods which lived in the Mississippian sub-period, a subdivision of the Carboniferous period. It contains the genera ''Pederpes'', '' Whatcheeria'', and possibly '' Ossinodus''. Fossils of a possible whatcheeriid have been found from the Red Hill locality of Pennsylvania. If these remains are from a whatcheeriid, they extend the range of the family into the Late Devonian and suggest that advanced tetrapods may have lived alongside primitive tetrapod ancestors like ''Hynerpeton'' and ''Densignathus''. They also imply that a very long ghost lineage of whatcheeriids lived through Romer's gap, a period during the Early Carboniferous conspicuously lacking in tetrapod remains. Classification Currently, using modern cladistic taxonomy, Whatcheeriidae is not placed in Amphibia Amphibians are four-limbed and ectothermic vertebrates of the class Amphibia. All living amphibians belong to the group Lissamphibia. They inhabit a wide va ...
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Ossinodus
''Ossinodus'' is an extinct genus of Crown group, stem tetrapod. Fossils have been found from the Ducabrook Formation in Queensland, Australia dating back to the middle Visean stage of the Early Carboniferous (Mississippian age, Mississippian). It was originally placed within the family (biology), family ''Whatcheeriidae'', but the absence of an intertemporal bone as suggested by a recent reconstruction of the skull based on fragmentary material may prove it to be stemward of all whatcheeriids. The oldest known pathological bone of a tetrapod, a fractured right radius, has been referred to ''Ossinodus''. History and discovery ''Ossinodus'' is one of the first large tetrapods discovered during the time known as Romer's gap, Romer’s Gap. In 1996, initial inspection of scraps and materials postulated three possible types of tetrapods. A premaxilla with a tusk was originally attributed to a Colosteidae, colosteid, then an ilium with a postiliac process linked it to an Anthracosaur ...
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Hynerpeton
''Hynerpeton'' ( ) is an extinct genus of early four-limbed vertebrate that lived in the rivers and ponds of Pennsylvania during the Late Devonian period, around 365 to 363 million years ago. The only known species of ''Hynerpeton'' is ''H. bassetti'', named after the describer's grandfather, city planner Edward Bassett. ''Hynerpeton'' is known for being the first Devonian four-limbed vertebrate discovered in the United States, as well as possibly being one of the first to have lost internal (fish-like) gills. This genus is known from few remains discovered at the Red Hill fossil site in Hyner, Pennsylvania. The most notable fossil is a large endochondral shoulder girdle consisting of the cleithrum, scapula, and coracoid (but not the interclavicle and clavicles), all connected into one shoulder bone. The inner surface of this shoulder bone possesses an array of depressions believed to have been attachment points for a unique set of powerful muscles around the chest. This may have ...
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Stegocephalia
Stegocephali (often spelled Stegocephalia) is a group containing all four-limbed vertebrates. It is equivalent to a broad definition of Tetrapoda: under this broad definition, the term "tetrapod" applies to any animal descended from the first vertebrate with limbs and toes, rather than fins. This includes both the modern lineage of limbed vertebrates (the crown group, including modern amphibians, mammals, reptiles and birds) as well as a portion of the stem group, limbed vertebrates that evolved prior to the origin of the crown group. Members of the tetrapod stem group include the earliest limbed tetrapodomorphs such as '' Ichthyostega'' and ''Acanthostega,'' which evolved in the Devonian Period long before any modern form of tetrapod. Many paleontologists prefer a stricter definition of Tetrapoda which applies solely to the crown group, excluding earlier types of limbed tetrapodomorphs. Stegocephali was re-established to replace the broad definition of Tetrapoda, resolving t ...
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Whatcheeria
''Whatcheeria'' is an extinct genus of early tetrapod from the Mississippian (Early Carboniferous) of Iowa. Fossils have been found in 340 million year old fissure fill deposits in the town of Delta. The type species, ''Whatcheeria deltae'' was named in 1995. It is classified within the family Whatcheeriidae, along with the closely related ''Pederpes'' and possibly ''Ossinodus.'' ''Whatcheeria'' is named after What Cheer, Iowa, the hometown of Pat McAdams, the geologist who discovered the first skeletons of the animal. The species is named after Delta, Iowa, the location where the fossils were uncovered. Description ''Whatcheeria'' possesses a mixture of both primitive and derived traits. It shares with earlier stem tetrapods a series of lateral lines across the skull, rows of teeth on the palate, and small Meckelian groove, Meckelian foramina across the surface of the lower jaw. It has a cleithrum, a bone in the pectoral girdle that extends from the scapula. The cleithrum o ...
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Ichthyostega BW
''Ichthyostega'' (from el, ἰχθῦς , 'fish' and el, στέγη , 'roof') is an extinct genus of limbed tetrapodomorphs from the Late Devonian of Greenland. It was among the earliest four-limbed vertebrates in the fossil record, and was one of the first with weight-bearing adaptations for terrestrial locomotion. ''Ichthyostega'' possessed lungs and limbs that helped it navigate through shallow water in swamps. Although ''Ichthyostega'' is often labelled a 'tetrapod' due to the possession of limbs and fingers, it evolved long before true crown group tetrapods, and could more accurately be referred to as a stegocephalian or stem tetrapod. Likewise, while undoubtedly of amphibian build and habit, it is not considered a true member of the group in the narrow sense, as the first modern amphibians (members of the group Lissamphibia) appeared in the Triassic Period. Until finds of other early stegocephalians and closely related fishes in the late 20th century, ''Ichthyostega'' stoo ...
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Pederpes
''Pederpes'' ("Peter's Foot") is an extinct genus of early Carboniferous tetrapod, dating from 348 to 347.6 Ma in the Tournaisian age (lower Mississippian). ''Pederpes'' contains one species, ''P. finneyae'', 1 m long. This most basal Carboniferous tetrapod had a large, somewhat triangular head, similar to that of later American sister-genus '' Whatcheeria'', from which it is distinguished by various skeletal features, such as a spike-like latissimus dorsi (an arm muscle) attachment on the humerus and several minor skull features. The feet had characteristics that distinguished it from the paddle-like feet of the Devonian Ichthyostegalia and resembled the feet of later, more terrestrially adapted Carboniferous forms. ''Pederpes'' is the earliest-known tetrapod to show the beginnings of terrestrial locomotion and despite the probable presence of a sixth digit on the forelimbs it was at least functionally pentadactyl. Discovery and classification ''Pederpes'' was discovered in 19 ...
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Ichthyostega
''Ichthyostega'' (from el, ἰχθῦς , 'fish' and el, στέγη , 'roof') is an extinct genus of limbed tetrapodomorphs from the Late Devonian of Greenland. It was among the earliest four-limbed vertebrates in the fossil record, and was one of the first with weight-bearing adaptations for terrestrial locomotion. ''Ichthyostega'' possessed lungs and limbs that helped it navigate through shallow water in swamps. Although ''Ichthyostega'' is often labelled a 'tetrapod' due to the possession of limbs and fingers, it evolved long before true crown group tetrapods, and could more accurately be referred to as a stegocephalian or stem tetrapod. Likewise, while undoubtedly of amphibian build and habit, it is not considered a true member of the group in the narrow sense, as the first modern amphibians (members of the group Lissamphibia) appeared in the Triassic Period. Until finds of other early stegocephalians and closely related fishes in the late 20th century, ''Ichthyostega'' st ...
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Tetrapod
Tetrapods (; ) are four-limbed vertebrate animals constituting the superclass Tetrapoda (). It includes extant and extinct amphibians, sauropsids ( reptiles, including dinosaurs and therefore birds) and synapsids (pelycosaurs, extinct therapsids and all extant mammals). Tetrapods evolved from a clade of primitive semiaquatic animals known as the Tetrapodomorpha which, in turn, evolved from ancient lobe-finned fish (sarcopterygians) around 390 million years ago in the Middle Devonian period; their forms were transitional between lobe-finned fishes and true four-limbed tetrapods. Limbed vertebrates (tetrapods in the broad sense of the word) are first known from Middle Devonian trackways, and body fossils became common near the end of the Late Devonian but these were all aquatic. The first crown-tetrapods (last common ancestors of extant tetrapods capable of terrestrial locomotion) appeared by the very early Carboniferous, 350 million years ago. The specific aquatic ancestors ...
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Elpistostegalia
Elpistostegalia or Panderichthyida is an order of prehistoric lobe-finned fishes which lived during the Middle Devonian to Late Devonian period (about 385 to 374 million years ago). They represent the advanced tetrapodomorph stock, the fishes more closely related to tetrapods than the osteolepiform fishes. The earliest elpistostegalians, combining fishlike and tetrapod-like characters, are sometimes called fishapods, a phrase coined for the advanced elpistostegalian ''Tiktaalik''. Through a strict cladistic view, the order includes the terrestrial tetrapods. Palaeobiology of the elpistostegalians A rise in global oxygen content allowed for the evolution of large, predatory fish that were able to exploit the shallow tidal areas and swamplands as top predators. Several groups evolved to fill these niches, the most successful were the elpistiostegalians. In such environments, they would have been challenged by periodic oxygen deficiency. In comparable modern aquatic environments l ...
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Elpistostege
''Elpistostege'' is an extinct genus of finned tetrapodomorphs that lived during the late Givetian to early Frasnian ages of the Late Devonian epoch. Its only known species, ''E. watsoni'', was first described in 1938 by the British palaeontologist Thomas Stanley Westoll, based on a single partial skull roof discovered at the Escuminac Formation in Quebec, Canada. In 2010, a complete specimen was found in the same formation, which was described by Richard Cloutier and colleagues in 2020. It reveals that the paired fins of ''Elpistostege'' contained bones homologous to the phalanges (digit bones) of modern tetrapods; it is the most basal tetrapodomorph known to possess these bones. At the same time, the fins were covered in scales and lepidotrichia (fin rays), which indicates that the origin of phalanges preceded the loss of fin rays, rather than the other way around. Relationships An analysis conducted by Swartz in 2012 found ''Elpistostege'' to be the sister taxon of ''Tiktaa ...
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Tiktaalik
''Tiktaalik'' (; Inuktitut ) is a monospecific genus of extinct sarcopterygian (lobe-finned fish) from the Late Devonian Period, about 375 Mya (million years ago), having many features akin to those of tetrapods (four-legged animals). It may have grown up to in length. Unearthed in Arctic Canada, ''Tiktaalik'' is a non-tetrapod member of Osteichthyes (bony fish), complete with scales and gills – but it has a triangular, flattened head and unusual, cleaver-shaped fins. Its fins have thin ray bones for paddling like most fish, but they also have sturdy interior bones that would have allowed Tiktaalik to prop itself up in shallow water and use its limbs for support as most four-legged animals do. Those fins and other mixed characteristics mark ''Tiktaalik'' as a crucial transition fossil, a link in evolution from swimming fish to four-legged vertebrates. This and similar animals might be the common ancestors of all vertebrate terrestrial fauna: amphibians, reptiles, birds, and m ...
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Tiktaalik BW
''Tiktaalik'' (; Inuktitut ) is a monospecific genus of extinct sarcopterygian (lobe-finned fish) from the Late Devonian Period, about 375 Mya (million years ago), having many features akin to those of tetrapods (four-legged animals). It may have grown up to in length. Unearthed in Arctic Canada, ''Tiktaalik'' is a non-tetrapod member of Osteichthyes (bony fish), complete with scales and gills – but it has a triangular, flattened head and unusual, cleaver-shaped fins. Its fins have thin ray bones for paddling like most fish, but they also have sturdy interior bones that would have allowed Tiktaalik to prop itself up in shallow water and use its limbs for support as most four-legged animals do. Those fins and other mixed characteristics mark ''Tiktaalik'' as a crucial transition fossil, a link in evolution from swimming fish to four-legged vertebrates. This and similar animals might be the common ancestors of all vertebrate terrestrial fauna: amphibians, reptiles, birds, and m ...
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