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Triclad
A planarian is one of the many flatworms of the traditional class Turbellaria. It usually describes free-living flatworms of the order Tricladida (triclads), although this common name is also used for a wide number of free-living platyhelminthes. Planaria are common to many parts of the world, living in both saltwater and freshwater ponds and rivers. Some species are terrestrial and are found under logs, in or on the soil, and on plants in humid areas. The triclads are characterized by triply branched intestine and anteriorly situated ovaries, next to the brain. Today the order Tricladida is split into three suborders, according to their phylogenetic relationships: Maricola, Cavernicola and Continenticola. Formerly, the Tricladida was split according to habitats: Maricola, which is marine; Paludicola which inhabits freshwater; and Terricola, which is land-dwelling. Planaria exhibit an extraordinary ability to regenerate lost body parts. For example, a planarian split len ...
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Dimarcusidae
Dimarcusidae is a family of triclads found mostly in freshwater habitats of caves, although at least one species, '' Rhodax evelinae'', occurs in surface waters. Currently the family contains only seven species distributed in five genera, although the total number of species is thought to be much higher. Description The morphological features uniting species of Dimarcusidae are related to the reproductive system. Their ovaries are located more posteriorly than in most triclads, which usually have them close to the brain. The penis in species of Dimarcusidae contains glandular elements and the common ovovitelline duct is perpendicular to the female genital duct. Taxonomy The family Dimarcusidae was erected in 1972 by Mitchell and Kawakatsu to include a new species, ''Dimarcus villalobosi''. However, the same species had been briefly described by Benazzi in the same year as '' Opisthobursa mexicana'' and this name had priority, with ''Dimarcus villalobosi'' becoming a junior ...
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Girardia Tigrina
''Girardia tigrina'' is a species of dugesiid native to the Americas.Ball, I. R.: A contribution to the phylogeny and biogeography of the freshwater triclads (Platyhelminthes: Turbellaria). Biology of the Turbellaria (Edited by: Riser NW and Morse MP). New York: McGraw-Hill New York 1974 , 339-401. It has been accidentally introduced into Europe and Japan.Sluys, R., Kawakatsu, M., Yamamoto, K., 2010. "Exotic freshwater planarians currently known from Japan". Belgian Journal of Zoology, 140 (Suppl.). p. 103-109 Description ''G. tigrina'' individuals are around 10 mm in length. They have a head with two broad and short auricles. The two eyes are in two pigment-free patches. The dorsal surface of the body has numerous pigment spots. Distribution and ecology Originally from the Americas, ''Girardia tigrina'' has become an invasive species in Europe and Japan. Since being first recorded in France in 1925, this species has been reported from many other European countries, includ ...
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Planaria Maculata
''Planaria'' is a genus of planarians in the family Planariidae. When an individual is cut into pieces, each piece has the ability to regenerate into a fully formed individual. Description Currently the genus ''Planaria'' is defined as freshwater triclads with oviducts that unite to form a common oviduct without embracing the bursa copulatrix and with an adenodactyl present in the male atrium. The testes occur along the whole body. Planaria originally have habitats in dark, murky water which results in such sensitivity (Paskin et al., 2014). They are also sensitive to other stimuli such as chemical gradients, vibration, magnetic and electric fields (Deochand et al., 2018). Their central nervous system includes the anterior (head, brain and eyes) and middle (abdominal trunk and pharynx) (Deochand et al., 2018). Diet The food of ''Planaria'' species includes freshwater gastropods, tubificid worms, and freshwater arthropods, such as isopods of the genus '' Asellus'' and chi ...
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Maricola
Maricola is a suborder of triclad flatworms including species that mainly inhabit salt water environments. However, some species are also known from freshwater or brackish waters.Ball, I. R. 1974. La Faune Terrestre de l'Ile de Saint Hélène: Turbellaria Tricladida. Ann. Mus. Roy. Afr. Centrale, in 8. Zool. Taxonomy and phylogeny History The Maricola group was first proposed by Hallez in 1892. He recognized three families: Otoplanida, Procerodida and Bdellourida. Two years later, in 1884, Hallez renamed these families as Otoplanidae, Procerodidae and Bdellouridae. In 1906 Böhmig classified the Maricola in two families and five subfamilies: Procerodidae (Euprocerodinae, Cercyrinae, Micropharynginae) and Bdellouridae (Uteriporinae, Eubdellourinae). In 1909 Wilhelmi wrote a monograph on the group in which five families were described: Procerodidae, Uteriporidae, Cercyridae, Bdellouridae, Micropharyngidae. Von Graff used the same classification in 1916. In 1989 Sluys recogniz ...
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Dugesiid
Dugesiidae is a family of freshwater planarians distributed worldwide (except Antarctica). The type genus is ''Dugesia'' Girard, 1850.Ball, I. R.: A contribution to the phylogeny and biogeography of the freshwater triclads (Platyhelminthes: Turbellaria). Biology of the Turbellaria (Edited by: Riser NW and Morse MP). New York: McGraw-Hill New York 1974, 339-401. Description All species of Dugesiidae live in freshwater environments and have a dorsoventrally flattened body. The head usually has a somewhat triangular shape and has two eyes (except for some subterranean eyeless species). The main differences between Dugesiidae and other freshwater planarians are related to the anatomy of the eyes and the copulatory apparatus. The eye cup in Dugesiidae is composed of several retinal cells, while in other freshwater planarians they are composed of a single cell. All freshwater planarians have an accessory organ called copulatory bursa or ''bursa copulatrix'', which is connected to the ...
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Terricola
Geoplanidae is a family of flatworms known commonly as land planarians or land flatworms. These flatworms are mainly predators of other invertebrates, which they hunt, attack and capture using physical force and the adhesive and digestive properties of their mucus. They lack water-retaining mechanisms and are therefore very sensitive to humidity variations of their environment. Because of their strict ecological requirements, some species have been proposed as indicators of the conservation state of their habitats. They are generally animals with low vagility (dispersal ability) and with very specific habitat requirements, so they can be also used to accurately determine the distribution of biogeographic realms. Today the fauna of these animals is being studied to select conservation priorities in the Atlantic rainforest in Brazil. At the other extreme, one species in this family, ''Platydemus manokwari'' has become an invasive species in both disturbed and wild habitats in the ...
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Schmidtea Mediterranea
''Schmidtea mediterranea'' is a freshwater triclad that lives in southern Europe and Tunisia. It is a model for regeneration, stem cells and development of tissues such as the brain and germline. Distribution ''Schmidtea mediterranea'' is found in some coastal areas and islands in the western Mediterranean (Catalonia, Menorca, Mallorca, Corsica, Sardinia, Sicily and Tunisia).Benazzi M, Baguñà J, Ballester R, Puccinelli I, Papa RD: Further Contribution to the Taxonomy of the ''Dugesia lugubris-polychroa Group'' with Description of ''Dugesia mediterranea'' n. sp. (Tricladida, Paludicola). Bolletino di zoologia 1975, 42(1):81-89.Ribas M: Cariologia, sistematica i biogeografia de les Planaries d'aigues dolces al Països Catalans. 1990.Baguñà J. Carranza S, Pala M, Ribera C, Giribet G, Arnedo M, Ribas M, Riutort M: From morphology and kariology to molecules. New methods for taxonomical identification of asexual populations of freshwater planarians. A tribute to Professor Mario Be ...
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Continenticola
Continenticola is a clade that includes the land planarians (Geoplanidae) and the freshwater triclads (Dendrocoelidae, Dugesiidae, Kenkiidae and Planariidae). Phylogeny On the basis of molecular evidences, Carranza and colleagues suggested in 1998 that the families of freshwater and land flatworms must be grouped together in a monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gr ... group that they coined Continenticola. This grouping was accepted by Ronald Sluys in 2009. Despite the molecular evidences, there are no morphological apomorphies supporting this clade. Phylogenetic supertree after Sluys et al., 2009: References {{Taxonbar, from=Q5165347 ...
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Cavernicola (suborder)
Cavernicola is a suborder of planarians found mostly in freshwater habitats of caves, although some species occur on the surface. Description The main morphological feature that defines the suborder Cavernicola is the placement of their ovaries. They are located more posteriorly than in other planarians, which usually have them close to the brain. The copulatory apparatus has a copulatory bursa in most species, although in some it is absent and is replaced by a connection to the intestine. Taxonomy The suborder Cavernicola was originally erected for the family Dimarcusidae Dimarcusidae is a family of triclads found mostly in freshwater habitats of caves, although at least one species, '' Rhodax evelinae'', occurs in surface waters. Currently the family contains only seven species distributed in five genera, althou ... due to its unusual morphological features that did not match those of the other planarian suborders. Recently, the discovery of another unusual cave spe ...
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Girardia Dorotocephala
''Girardia dorotocephala'' is a species of dugesiid triclad native to North America. It has been accidentally introduced in Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ....Sluys, R., Kawakatsu, M., Yamamoto, K. (2010). "Exotic freshwater planarians currently known from Japan". ''Belgian Journal of Zoology'', 140 (Suppl.). p. 103-109 ''Girardia dorotocephala'' is cannibalistic, which led to its usage in memory transfer experiments. References Dugesiidae {{flatworm-stub ...
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Endoderm
Endoderm is the innermost of the three primary germ layers in the very early embryo. The other two layers are the ectoderm (outside layer) and mesoderm (middle layer). Cells migrating inward along the archenteron form the inner layer of the gastrula, which develops into the endoderm. The endoderm consists at first of flattened cells, which subsequently become columnar. It forms the epithelial lining of multiple systems. In plant biology, endoderm corresponds to the innermost part of the cortex (bark) in young shoots and young roots often consisting of a single cell layer. As the plant becomes older, more endoderm will lignify. Production The following chart shows the tissues produced by the endoderm. The embryonic endoderm develops into the interior linings of two tubes in the body, the digestive and respiratory tube. Liver and pancreas cells are believed to derive from a common precursor. In humans, the endoderm can differentiate into distinguishable organs after 5 wee ...
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Dermis
The dermis or corium is a layer of skin between the epidermis (with which it makes up the cutis) and subcutaneous tissues, that primarily consists of dense irregular connective tissue and cushions the body from stress and strain. It is divided into two layers, the superficial area adjacent to the epidermis called the papillary region and a deep thicker area known as the reticular dermis.James, William; Berger, Timothy; Elston, Dirk (2005). ''Andrews' Diseases of the Skin: Clinical Dermatology'' (10th ed.). Saunders. Pages 1, 11–12. . The dermis is tightly connected to the epidermis through a basement membrane. Structural components of the dermis are collagen, elastic fibers, and extrafibrillar matrix.Marks, James G; Miller, Jeffery (2006). ''Lookingbill and Marks' Principles of Dermatology'' (4th ed.). Elsevier Inc. Page 8–9. . It also contains mechanoreceptors that provide the sense of touch and thermoreceptors that provide the sense of heat. In addition, hair foll ...
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