Therizinosaurid
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Therizinosaurid
Therizinosauridae (meaning 'scythe lizards')Translated paper
is a family of derived (advanced) dinosaurs whose fossil remains have been found in mostly boundary. Even though representative fossils have only been found throughout and North America, the range of Therizino ...
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Therizinosauroid
Therizinosaurs (once called segnosaurs) were large herbivorous theropod dinosaurs whose fossils have been found across the Early to Late Cretaceous deposits in Asia and North America. Various features of the forelimbs, skull and pelvis unite these finds as both theropods and maniraptorans, close relatives to birds. The name of the representative genus, ''Therizinosaurus'', is derived from the Greek (, 'to reap' or 'scythe')Translated paper
and (, 'lizard'). The older representative, ''Segnosaurus'', is derived from the ('slow') and the Greek .


History of research


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Therizinosaurus Cheloniformis
''Therizinosaurus'' (; meaning 'scythe lizard') is a genus of very large therizinosaurid that lived in Asia during the Late Cretaceous period in what is now the Nemegt Formation around 70 million years ago. It contains a single species, ''Therizinosaurus cheloniformis''. The first remains of ''Therizinosaurus'' were found in 1948 by a Mongolian field expedition at the Gobi Desert and later described by Evgeny Maleev in 1954. The genus is only known from a few bones, including gigantic manual unguals (claw bones), from which it gets its name, and additional findings comprising fore and hindlimb elements that have been discovered during the 1960s–1980s. ''Therizinosaurus'' was a colossal therizinosaurid that could grow up to long and tall, and weigh possibly over . Like other therizinosaurids, it would have been a slow-moving, long-necked, high browser equipped with a rhamphotheca (horny beak) and a wide torso for food processing. Its forelimbs were particularly robust and had t ...
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Therizinosaurus
''Therizinosaurus'' (; meaning 'scythe lizard') is a genus of very large therizinosaurid that lived in Asia during the Late Cretaceous period in what is now the Nemegt Formation around 70 million years ago. It contains a single species, ''Therizinosaurus cheloniformis''. The first remains of ''Therizinosaurus'' were found in 1948 by a Mongolian field expedition at the Gobi Desert and later described by Evgeny Maleev in 1954. The genus is only known from a few bones, including gigantic manual unguals (claw bones), from which it gets its name, and additional findings comprising fore and hindlimb elements that have been discovered during the 1960s–1980s. ''Therizinosaurus'' was a colossal therizinosaurid that could grow up to long and tall, and weigh possibly over . Like other therizinosaurids, it would have been a slow-moving, long-necked, high browser equipped with a rhamphotheca (horny beak) and a wide torso for food processing. Its forelimbs were particularly robust and had t ...
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Segnosaurus
''Segnosaurus'' is a genus of therizinosaurid dinosaur that lived in what is now southeastern Mongolia during the Late Cretaceous, about 102–86 million years ago. Multiple incomplete but well-preserved specimens were discovered in the Gobi Desert in the 1970s, and in 1979 the genus and species ''Segnosaurus galbinensis'' were named. The generic name ''Segnosaurus'' means "slow lizard" and the specific name ''galbinensis'' refers to the Galbin region. The known material of this dinosaur includes the lower jaw, neck and tail vertebrae, the pelvis, shoulder girdle, and limb bones. Parts of the specimens have gone missing or become damaged since they were collected. ''Segnosaurus'' was a large-bodied therizinosaur that is estimated to have been about long and to have weighed about . It would have been bipedal, with the trunk of its body tilted upwards. The head was small with a beak at the tip of the jaws, and the neck was long and slender. The lower jaw was down-turned at t ...
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Erlikosaurus
''Erlikosaurus'' (meaning "Erlik's lizard") is a genus of therizinosaurid that lived in Asia during the Late Cretaceous period. The fossils, a skull and some post-cranial fragments, were found in the Bayan Shireh Formation of Mongolia in 1972, dating to around 96 million and 89 million years ago. These remains were later described by Altangerel Perle and Rinchen Barsbold in 1980, naming the new genus and species ''Erlikosaurus andrewsi''. It represents the second therizinosaur taxon from this formation (alongside ''Enigmosaurus'' and ''Segnosaurus'') with the most complete skull among members of this peculiar family of dinosaurs. In contrast to most therizinosaurids, ''Erlikosaurus'' was a small member reaching nearly in length and in mass. It had a well-developed beak at the snout tip and toothed jaws that were used for its herbivorous diet. The feet ended in four toes with the first one articulated to the ankle—in contrast to the vestigial first toe of most theropods. Like o ...
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Nothronychus
''Nothronychus'' (meaning "slothful claw") is a genus of therizinosaurid theropod dinosaurs that lived in North America during the Late Cretaceous period. The type species, ''Nothronychus mckinleyi'', was described by James Kirkland and Douglas G. Wolfe in 2001. It was recovered near New Mexico's border with Arizona, in an area known as the Zuni Basin, from rocks assigned to the Moreno Hill Formation, dating to the late Cretaceous period (mid-Turonian stage), around 91 million years ago. A second specimen, described in 2009 as a second species, ''Nothronychus graffami'', was found in the Tropic Shale of Utah, dating to the early Turonian, between one million and a half million years older than ''N. mckinleyi''. ''Nothronychus'' were bulky herbivorous theropods with wide, sloth-like hip (resembling that of the non-related ornithischians), four-toed feet with all four toes facing forward, elongated necks and prominent arms with sharp claws. Both species were similar in dimensions, ...
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Nanshiungosaurus
''Nanshiungosaurus'' (meaning "Nanxiong's lizard") is a genus of therizinosaurid that lived in what is now Asia during the Late Cretaceous of South China. The type species, ''Nanshiungosaurus brevispinus'', was first discovered in 1974 and described in 1979 by Dong Zhiming. It is represented by a single specimen preserving most of the cervical and dorsal vertebrae with the pelvis. A supposed and unlikely second species, ''"Nanshiungosaurus" bohlini'', was found in 1992 and described in 1997. It is also represented by vertebrae but this species however, differs in geological age and lacks authentic characteristics compared to the type, making its affinity to the genus unsupported. It was a large-sized therizinosaurid that is estimated to have cover nearly in length and to have weighed about . ''Nanshiungosaurus'' had a very pneumatized vertebral column with the posterior cervical vertebrae of the long neck being unusually robust and slightly more elongated than the dorsals. It wa ...
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Evgeny Maleev
Evgeny Aleksandrovich Maleev (, ; 25 February 1915 – 12 April 1966) was a Soviet and Russian paleontologist who did most of his research on reptiles and Asian fossils, such as the naming of the ankylosaur ''Talarurus'' and theropods ''Tarbosaurus'' and ''Therizinosaurus'' along with the family Therizinosauridae. Biography Evgeny Aleksandrovich Maleev was born on February 25, 1915, in the Russian Empire. After being a member on the Eastern Front (World War II) he started his scientific career at the Moscow State University. In 1947 he graduated from the Faculty of Biology, also in that year, he started to work at the Paleontological Institute, Russian Academy of Sciences. In 1950 Maleev presented his PhD thesis, "Morphofunctional analysis of the occipital region of the skull and skeleton of the neck of mammals", being awarded the Candidate of Sciences. Later on, he worked as the deputy manager from 1956 to 1962. He also was the leader of an expedition conducted in 1962 to In ...
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Suzhousaurus
''Suzhousaurus'' (meaning "Suzhou lizard") is a genus of large therizinosauroid dinosaur from the Early Cretaceous of China. The genus is known from two specimens discovered on the Xiagou Formation and Zhonggou Formation—which are situated in the Xinminbao Group. These findings were made during field-works in 1999 and 2004. Though ''Suzhousaurus'' is known from these two specimens, an earlier named and described therizinosauroid from the adjacent basin, ''"Nanshiungosaurus" bohlini'', may be synonymous with the former. However, ''Suzhousaurus'' can not be compared to this species due to non-overlapping material and the loss of the same. Moreover, this synonymy will result in ''Suzhousaurus bohlini'' with ''"N". bohlini'' having priority. ''Suzhousaurus'' was an unusually large Early Cretaceous therizinosauroid reaching lengths of and nearly in weight. The upper arm (humerus) was very distinct from therizinosaurids, making ''Suzhousaurus'' to be considered as a therizinosauro ...
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Erliansaurus
''Erliansaurus'' (meaning "Erlian lizard") is a genus of therizinosaur theropod dinosaur that lived in Asia during the Cenomanian stage of the Late Cretaceous period in what is now Nei Mongol, Iren Dabasu Formation. Discovery and naming The remains of ''Erliansaurus'' were found near Sanhangobi in Inner Mongolia in 1999. The type species, ''Erliansaurus bellamanus'', was described by Xu Xing, Zhang Xiaohong, Paul Sereno, Zhao Xijin, Kuang Xuewen, Han Jun and Tan Lin in 2002. The generic name refers to the town of Erlian and the specific name is derived from Latin ''bellus'', "beautiful", and ''manus'', "hand", in reference to the exquisite preservation of the forelimb. The holotype, LH V0002, was uncovered in the Iren Dabasu Formation dating from the Cenomanian stage. It consists of a partial skeleton belonging to a subadult individual. It includes two cervical, one dorsal and two caudal vertebrae; a right scapula; a left forelimb only lacking the carpus; a partial right ...
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Paralitherizinosaurus
''Paralitherizinosaurus'' is an extinct genus of therizinosaurid dinosaur from the Late Cretaceous Osoushinai Formation of Hokkaido, Japan. The genus contains a single species, ''P. japonicus'', known from a partial right hand and cervical vertebra. ''Paralitherizinosaurus'' represents the youngest therizinosaur known from Japan. Discovery and naming The ''Paralitherizinosaurus'' holotype specimen, NMV-52, was discovered in September 2000 in layers of the Osoushinai Formation (Yezo Group) in Nakagawa, Hokkaido, Japan, which dates to the early Campanian age of the late Cretaceous period. The specimen consists of a partial cervical vertebra and the metacarpal I, proximal ends of unguals I and II, and nearly complete ungual III of the right hand. Some of these remains were first described by Murakami ''et al''. in 2008 as belonging to an indeterminate genus of maniraptoran theropod, possibly with therizinosauroid affinities. In 2022, Kobayashi ''et al''. described ''Paralitheriz ...
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Neimongosaurus
''Neimongosaurus'' (meaning "Nei Mongol lizard") is a genus of herbivorous therizinosaur theropod dinosaur that lived in Asia during the Cenomanian stage of the Late Cretaceous period in what is now the Iren Dabasu Formation. Discovery and naming ''Neimongosaurus'' is a therizinosauroid which is known from the holotype LH V0001, consisting of a partially preserved braincase; the front of the right lower jaw; a nearly complete axial column compromising 15 cervical (including the axis), 4 dorsal and 22 caudal vertebrae; a furcula; both scapulocoracoids; both humeri; left radius; fragmented ilia; both femora; both tibiae; left tarsals and a virtually complete and articulated left pes. A second specimen, LH V0008, consisting of a sacrum composed by 6 sacral vertebrae and both ilia, was assigned as the paratype. The specimens were collected in 1999 at Sanhangobi in Inner Mongolia from the Iren Dabasu Formation, dating from the Cenomanian stage. Based on these remains, the type sp ...
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