Testudinata
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Testudinata
Testudinata is the group of all tetrapods with a true turtle shell. It includes both modern turtles (Testudines) and many of their extinct, shelled relatives (stem-turtles). Though it was first coined as the group containing turtles by Jacob Theodor Klein in 1760, it was first defined in the modern sense by Joyce and colleagues in 2004. Testudinata does not include the primitive stem-turtle '' Odontochelys'', which only had the bottom half of a shell. A recent phylogenetic tree of Testudinata included Angolachelonia Angolachelonia is a clade of extinct turtles from the Late Jurassic to Paleogene of Eurasia. The group is defined as all taxa derived from the ancestor of the type genus ''Angolachelys'' and '' Solnhofia'', a definition that could potentially e ... and Testudines as sister-taxa and subgroups of Testudinata. Classification The cladogram below follows an analysis by Jérémy Anquetin in 2012. References {{Taxonbar, from=Q16114972 Reptile taxonomy Nori ...
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Chuannanchelys
Testudinata is the group of all tetrapods with a true turtle shell. It includes both modern turtles (Testudines) and many of their extinct, shelled relatives (stem-turtles). Though it was first coined as the group containing turtles by Jacob Theodor Klein Jacob Theodor Klein (nickname ''Plinius Gedanensium''; 15 August 1685 – 27 February 1759) was a German jurist, historian, botanist, zoologist, mathematician and diplomat in service of Polish King August II the Strong. Life Klein was born on 15 ... in 1760, it was first defined in the modern sense by Joyce and colleagues in 2004. Testudinata does not include the primitive stem-turtle '' Odontochelys'', which only had the bottom half of a shell. A recent phylogenetic tree of Testudinata included Angolachelonia and Testudines as sister-taxa and subgroups of Testudinata. Classification The cladogram below follows an analysis by Jérémy Anquetin in 2012. References {{Taxonbar, from=Q16114972 Reptile taxonomy Nor ...
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Kayentachelys
''Kayentachelys'' ("Kayenta turtle") is an extinct genus of turtle known only from the "silty facies" of the Lower Jurassic Kayenta Formation in northeastern Arizona on the lands of the Navajo Nation. History of discovery and significance The earliest mention of turtles from the Kayenta Formation in the literature is within the description of the ornithischian dinosaur '' Scutellosaurus'' by Colbert (1981). These first specimens were collected during the 1970s and early 1980s by field parties from the Museum of Northern Arizona (MNA) and the Museum of Comparative Zoology (MCZ) at Harvard University. Specimens from these field excursions were used to establish the taxon ''Kayentachelys'' in 1987, and it is an MNA specimen (MNA V1558) which was designated as the holotype of the taxon. During the early 1980s, addition turtle specimens from the Kayenta Formation were collected by field parties from the University of California Museum of Paleontology (UCMP), which were repor ...
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Eileanchelys
''Eileanchelys'' is an extinct genus of primitive turtle from the Middle Jurassic ( Bathonian) period some 164 million years ago of Britain. Only one species is recorded, ''Eileanchelys waldmani''. It is the best-represented turtle from the Middle Jurassic, because of the amount of specimens that can be assigned to it. The turtle is also one of the oldest turtles ever found to be aquatic, and might represent a milestone in turtle evolution. Discovery and naming In 2009, two specimens from the National Museums of Scotland, Edinburgh, including NMS G 2004.31.15 and NMS G 2004.31.16a–f, were found to differ from other turtles from the area and age. It was decided that they represented a new genus and species, and were named ''Eileanchelys waldmani'' by Jérémy Anquetin and his colleagues. The specimen NMS G 2004.31.15 was designated as the holotype, and NMS G 2004.31.16a-f, the paratypes. NMS G 2004.31.16 includes at least five, and possibly six, different individuals, al ...
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Proterochersis
''Proterochersis'' is an extinct genus of turtle from the Late Triassic period (Norian The Norian is a division of the Triassic Period. It has the rank of an age (geochronology) or stage (chronostratigraphy). It lasted from ~227 to million years ago. It was preceded by the Carnian and succeeded by the Rhaetian. Stratigraphic defi ... stage) of Europe. It is known from a large number of fossils uncovered in Germany and Poland.Fraas E (1913)"''Proterochersis'', eine pleurodire Schildkröte aus dem Keuper" ''Jahreshefte des Vereins für Vaterländische Naturkunde in Württemberg'' 69: p. 13–30Szczygielski T & Sulej T (2016). "Revision of the Triassic European turtles ''Proterochersis'' and ''Murrhardtia'' (Reptilia, Testudinata, Proterochersidae), with the description of new taxa from Poland and Germany". ''Zoological Journal of the Linnean Society'' 177(2): pp. 395–427. Szczygielski1 T, Słowiak J & Dróżdż1 D (2018). "Shell variability in the stem turtles ''Proterocher ...
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Mesochelydia
Mesochelydia (from Greek ''mesos'' "middle" and ''chelys'' "turtle") is a clade within Pantestudines, more inclusive than Perichelydia, but less than Testudinata. The clade is known from the Early Jurassic to the Present, and contains all Jurassic representatives of Testudinata aside from ''Australochelys.'' The ancestral condition for Mesochelydia is thought to be aquatic, as opposed to terrestrial for Testudinata. They are distinguished from more basal testudinatans by the presence of the following characters: strap like pectoral girdle, supramarginals absent, reduced posterior entoplastral process, eleven pairs of peripherals, elongate processus interfenestralis, paired basioccipital tubercles, fully formed cavum tympani and antrum postoticum, single vomer, confluent external nares, lacrimals and supratemporals absent. Taxonomy * ''Condorchelys'' Sterli, 2008 Cañadón Asfalto Formation, Middle Jurassic, Argentina * ''Eileanchelys'' Anquetin et al. 2009 Kilmaluag Formation, ...
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Chinlechelys
''Chinlechelys'' ( meaning ''Chinle turtle'') is an extinct genus of stem-turtle belonging to Testudinata. It lived in the Norian age of the Late Triassic and is the oldest turtle known from North America. Among turtles it is unique, mostly because of its very thin shell. The type and only species, ''C. tenertesta'', was named and described with the genus by Walter G. Joyce ''et al.'' in 2009. It was probably terrestrial, and was found by Joyce ''et al.'' to be closely related to ''Proganochelys'', another terrestrial testudinatan. Discovery and naming ''Chinlechelys'' is known from only one assigned specimen, the holotype NMMNH P-16697 (New Mexico Museum of Natural History and Science) recovered from the Norian Bull Canyon Formation (New Mexico, United States). The skeleton consists of the middle of the carapace, the left hypoplastron, rear ribs, a neck spine, isolated osteoderms, and portions of the bridge. The materials were collected separately over several years, but exhib ...
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Testudines
Turtles are an order (biology), order of reptiles known as Testudines, characterized by a special turtle shell, shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Cryptodira (hidden necked turtles), which differ in the way the head retracts. There are 360 living and recently extinct species of turtles, including land-dwelling tortoises and freshwater terrapins. They are found on most continents, some islands and, in the case of sea turtles, much of the ocean. Like other Amniote, amniotes (reptiles, birds, and mammals) they breathe air and do not lay eggs underwater, although many species live in or around water. Turtle shells are made mostly of bone; the upper part is the domed Turtle shell#Carapace, carapace, while the underside is the flatter plastron or belly-plate. Its outer surface is covered in scale (anatomy), scales made of keratin, the material of hair, horns, and claws. The carapace bon ...
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Proganochelys Quenstedti
''Proganochelys'' is an extinct, primitive stem-turtle that has been hypothesized to be the sister taxon to all other turtles creating a monophyletic group, the ''Casichelydia''. ''Proganochelys'' was named by Georg Baur in 1887 as the oldest turtle in existence at the time. The name ''Proganochelys'' comes from the Greek word ''ganos'' meaning 'brightness', combined with prefix ''pro'', 'before', and Greek base ''chelys'' meaning 'turtle'. ''Proganochelys'' is believed to have been around 1 meter in size and herbivorous in nature. ''Proganochelys'' was known as the most primitive stem-turtle for over a century, until the novel discovery of ''Odontochelys'' in 2008. ''Odontochelys'' and ''Proganochelys'' share unique primitive features that are not found in ''Casichelydia'', such as teeth on the pterygoid and vomer and a plate-like coracoid. ''Proganochelys'' is the oldest stem-turtle species with a complete shell discovered to date, known from fossils found in Germany, Switzerla ...
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Odontochelys
''Odontochelys semitestacea'' (meaning "toothed turtle with a half-shell") is a Late Triassic relative of turtles. Before ''Pappochelys'' was discovered and ''Eunotosaurus'' was redescribed, ''Odontochelys'' was considered the oldest undisputed member of Pantestudines (i.e. a stem-turtle). It is the only known species in the genus ''Odontochelys'' and the family Odontochelyidae. Discovery ''Odontochelys semitestacea'' was first described from three 220-million-year-old specimens excavated in Triassic deposits in Guizhou, China. The locale of its discovery at one time was the Nanpanjiang Trough basin, a shallow marine environment surrounded on three sides by land. These deposits preserve an ecosystem known as the Guanling biota, which was dominated by marine reptiles. Description ''Odontochelys'' differed grossly from modern turtles. Modern turtles have a horny beak without teeth in their mouth. In contrast, ''Odontochelys'' fossils were found to have had teeth embedded in their ...
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