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Titanosauriform
Macronaria is a clade of sauropod dinosaurs. Macronarians are named after the large diameter of the nasal opening of their skull, known as the external naris, which exceeded the size of the orbit, the skull opening where the eye is located (hence ''macro''- meaning large, and –''naria'' meaning nose). Fossil evidence suggests that macronarian dinosaurs lived from the Middle Jurassic (Bathonian) through the Late Cretaceous (Maastrichtian). Macronarians have been found globally, including discoveries in Argentina, the United States, Portugal, China, and Tanzania. Like other sauropods, they are known to have inhabited primarily terrestrial areas, and little evidence exists to suggest that they spent much time in coastal environments. Macronarians are diagnosed through their distinct characters on their skulls, as well as appendicular and vertebral characters. Macronaria is composed of several subclades and families notably including Camarasauridae and Titanosauriformes, among seve ...
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Brachiosauridae
The Brachiosauridae ("arm lizards", from Greek ''brachion'' (βραχίων) = "arm" and ''sauros'' = "lizard") are a family or clade of herbivorous, quadrupedal sauropod dinosaurs. Brachiosaurids had long necks that enabled them to access the leaves of tall trees that other sauropods would have been unable to reach. In addition, they possessed thick spoon-shaped teeth which helped them to consume tough plants more efficiently than other sauropods. They have also been characterized by a few unique traits or synapomorphies; dorsal vertebrae with 'rod-like' transverse processes and an ischium with an abbreviated pubic peduncle. ''Brachiosaurus'' is one of the best-known members of the Brachiosauridae, and was once thought to be the largest land animal to ever live. Brachiosaurids thrived in the regions which are now North and South America, Africa, Europe, and Asia. They first appear in the fossil record in the Late Jurassic Period (possibly even earlier in the Middle Jurassic) a ...
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"Brachiosaurus" Nougaredi
''Brachiosaurus'' () is a genus of sauropod dinosaur that lived in North America during the Late Jurassic, about 154to 150million years ago. It was first described by American paleontologist Elmer S. Riggs in 1903 from fossils found in the Colorado River valley in western Colorado, United States. Riggs named the dinosaur ''Brachiosaurus altithorax''; the generic name is Greek for "arm lizard", in reference to its proportionately long arms, and the specific name means "deep chest". ''Brachiosaurus'' is estimated to have been between 18 and 22 meters (59 and 72ft) long; body mass estimates of the subadult holotype specimen range from 28.3 to 46.9 metric tons (31.2 and 51.7 short tons). It had a disproportionately long neck, small skull, and large overall size, all of which are typical for sauropods. Atypically, ''Brachiosaurus'' had longer forelimbs than hindlimbs, which resulted in a steeply inclined trunk, and a proportionally shorter tail. ''Brachiosaurus'' is the namesak ...
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Somphospondyli
Somphospondyli is an extinct clade of titanosauriform sauropods that lived from the Late Jurassic until the end of the Late Cretaceous, comprising all titanosauriforms more closely related to Titanosauria proper than Brachiosauridae. The remains of somphospondylans have been discovered in all six continents. Classification The group has officially been defined under the PhyloCode as the largest clade containing ''Saltasaurus loricatus'', but not ''Giraffatitan brancai.'' Features found as diagnostic of this clade by Mannion ''et al.'' (2013) include the possession of at least 15 cervical vertebrae; a bevelled radius bone end; sacral vertebrae with camellate internal texture; convex posterior articular surfaces of middle to posterior caudal vertebrae; biconvex distal caudal vertebrae; humerus anterolateral corner "squared"; among multiple others. The following cladogram depicts the reference phylogeny used to defined Somphospondlyi under the PhyloCode The ''International Code ...
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Fukuititan
''Fukuititan'' (meaning "Fukui giant") is a genus of titanosauriform sauropod dinosaur that lived in the Early Cretaceous (either Barremian or Aptian age) in what is now Japan. It is known from FPDM-V8468, the associated partial skeleton of a single individual, recovered from the Kitadani Dinosaur Quarry (Kitadani Formation) of the Tetori Group, at Katsuyama City. The type species, ''Fukuititan nipponensis'', was described in 2010 by Japanese scientists Yoichi Azuma and Masateru Shibata of the Fukui Prefectural Dinosaur Museum The , located in Katsuyama, Fukui, Japan, is one of the leading dinosaur museums in Asia that is renowned for its exhibits of fossil specimens of dinosaurs and paleontological research. It is sited in the Nagaoyama Park ( Katsuyama Dinosaur For .... The discovery sheds light on Japanese titanosauriforms, which are poorly known in the region. References Early Cretaceous dinosaurs of Asia Fossil taxa described in 2010 Macronarians Fossils of ...
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Fushanosaurus
''Fushanosaurus'' (meaning "Fushan lizard", after the Fushan Museum where its remains are stored) is a genus of basal titanosauriform dinosaur from the Shishugou Formation from Xinjiang Province in China. The type and only species is ''Fushanosaurus qitaiensis''. It is solely known from the holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ... specimen (FH000101), which consists of a complete right femur. Using "Huanghetitan" ''ruyangensis'' and '' Daxiatitan'' as a basis, ''Fushanosaurus'' was estimated to have been approximately long, which would then make it one of the longest known dinosaurs. It was published in the journal Global Geology. References Macronarians Callovian life Oxfordian life Middle Jurassic dinosaurs of Asia Late Jurassic dinosaurs of A ...
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Pukyongosaurus
''Pukyongosaurus'' (meaning "Pukyong lizard", after the Pukyong National University) is a genus of titanosauriform dinosaur that lived in South Korea during the Early Cretaceous period (Aptian - Albian). It may have been closely related to ''Euhelopus'', and is known from a series of vertebrae in the neck and back. The characteristics that were originally used to distinguish this genus have been criticized as being either widespread or too poorly preserved to evaluate, rendering the genus an indeterminate nomen dubium among titanosauriforms. The 2022 study noted that ''Pukyongosaurus'' is probably a somphospondylan. Discovery In 2001, several fragments of a sauropod skeleton were discovered in the Hasandong Formation in Hadong County, South Korea South Korea, officially the Republic of Korea (ROK), is a country in East Asia, constituting the southern part of the Korea, Korean Peninsula and sharing a Korean Demilitarized Zone, land border with North Korea. Its western borde ...
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Brohisaurus
''Brohisaurus'' is a genus of titanosauriform sauropod dinosaur from the Late Jurassic, based on largely indeterminate fragments of some ribs, vertebrae, and limb bones. The type and only species, ''B. kirthari'', was described by M. Sadiq Malkani in 2003 and it is currently the only valid genus of dinosaur known from Pakistan to date. The genus name means "Baluchi lizard" and refers to the Baluchi people who live in the area where it was found. The species name refers to the Kirthar Mountains. The fossils were discovered in the lowest portion of the Kimmeridgian Sembar Formation from the Kirthar foldbelt in Pakistan. Description ''Baluchisaurus'', like all sauropod dinosaurs, would have been a large-bodied, long-necked herbivore. Its femur was only 12 cm across. The 15 to 20 meter long titanosauriform '' Phuwiangosaurus'', by contrast, had a femur 20 cm in diameter. Classification ''Brohisaurus'' was originally described as a titanosaur. Malkani suggested it was ...
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Huanghetitan
''Huanghetitan'' (meaning "Yellow River titan"), is a genus of sauropod dinosaur from the early Cretaceous Period. It was a basal titanosauriform which lived in what is now Gansu, China. History The type species, ''Huanghetitan liujiaxiaensis'', was described by You ''et al.'' in 2006. It is known from fragmentary materials including two caudal vertebrae, an almost complete sacrum, rib fragments, and the left shoulder girdle, and was discovered in the eastern part of the Lanzhou Basin (Hekou Group) in the Gansu Province in 2004. A second species, ''H. ruyangensis'', was described in 2007 from the Aptian-Albian Haoling Formation of Ruyang County, China (Henan Province). A recent cladistic analysis has found that this species is unlikely to be closely related to ''H. liujiaxiaensis'' and requires a new genus name. Description ''H. liujiaxiaensis'' is a relatively small sauropod, measuring long and weighing . ''H. ruyangensis'' is known from a partial vertebral column ...
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Sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their body), and four thick, pillar-like legs. They are notable for the enormous sizes attained by some species, and the group includes the largest animals to have ever lived on land. Well-known genera include '' Brachiosaurus'', '' Diplodocus'', ''Apatosaurus'' and '' Brontosaurus''. The oldest known unequivocal sauropod dinosaurs are known from the Early Jurassic. '' Isanosaurus'' and '' Antetonitrus'' were originally described as Triassic sauropods, but their age, and in the case of ''Antetonitrus'' also its sauropod status, were subsequently questioned. Sauropod-like sauropodomorph tracks from the Fleming Fjord Formation (Greenland) might, however, indicate the occurrence of the group in the Late Triassic. By the Late Jurassic (150 mil ...
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Janenschia
''Janenschia'' (named after Werner Janensch) is a large herbivorous sauropod dinosaur from the Late Jurassic Tendaguru Formation of Lindi Region, Tanzania around 155 million years ago. Discovery and naming ''Janenschia'' has had a convoluted nomenclatural history. In 1907, Eberhard Fraas at "site P", nine hundred metres to the southeast of Tendaguru Hill, discovered two skeletons of gigantic sauropods. They were designated as "Skeleton A" and "Skeleton B". The fossils were transported to the collection of the ''Stuttgarter Naturaliensammlung'' in Stuttgart, Germany. Fraas in 1908 decided to name both skeletons as different species of one genus, ''Gigantosaurus''. Skeleton A became ''Gigantosaurus africanus'' and skeleton B became ''Gigantosaurus robustus''. The latter species was based on the holotype partial skeleton SMNS 12144, consisting of a right hindlimb. The specific name was inspired by the heavy build of the animal. While doing so, Fraas knew full well that the name ...
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Astrodon
''Astrodon'' (aster: star, odon: tooth) is a genus of large herbivorous sauropod dinosaur, measuring in length, in height and in body mass. It lived in what is now the eastern United States during the Early Cretaceous period, and fossils have been found in the Arundel Formation, which has been dated through palynomorphs to the Albian about 112 to 110 million years ago. Discovery and species Two dinosaur teeth were received in late November 1858 by chemist Philip Thomas Tyson from John D. Latchford. They had been found in Latchford's open iron ore pit in the Arundel Formation at Swampoodle near Muirkirk in Prince George's County, Maryland. Tyson let them be studied by the dentist Christopher Johnston, professor at the Baltimore Dental College, who cut one tooth in half and thereby discovered a characteristic star-formed cross-section. Johnston named ''Astrodon'' in 1859. However, he did not attach a specific epithet, so Joseph Leidy is credited with naming ''Astrodon joh ...
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Ornithopsis
''Ornithopsis'' (meaning "bird-likeness") was a medium-sized Early Cretaceous sauropod dinosaur, from England. The type species, which is the only species seen as valid today, is ''O. hulkei''. History of discovery Gideon Algernon Mantell described many fossils that had been previously collected from the Tilgate Forest of the Early Cretaceous Wealden Formation in his 1833 paper on the geology of southeast England, including a bone he considered to be the of ''Iguanodon'', otherwise only known definitively from teeth that had been found in the area since 1822. The bone was redescribed by Richard Owen in 1854, who reaffirmed its referral as a quadrate of ''Iguanodon'', but also suggested it could be the same bone of ''Streptospondylus'' or ''Cetiosaurus'' as it was not directly associated with the characteristic teeth of ''Iguanodon''. This specimen is stored as British Museum of Natural History R2239, having been purchased from Mantell in 1838. BMNH R28632, a similar bone to BMN ...
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