Steppe Lion
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Steppe Lion
''Panthera spelaea'', also known as the Eurasian cave lion, European cave lion or steppe lion, is an extinct ''Panthera'' species that most likely evolved in Europe after the third Cromerian interglacial stage, less than 600,000 years ago. Phylogenetic analysis of fossil bone samples revealed that it was highly distinct and genetically isolated from the modern lion (''Panthera leo'') occurring in Africa and Asia. Analysis of morphological differences and mitochondrial data support the taxonomic recognition of ''Panthera spelaea'' as a distinct species that genetically diverged from the lion about . Nuclear genomic evidence shows a more recent split approximately 500,000 years ago, with no subsequent interbreeding with the ancestors of the modern lion. The oldest known bone fragments were excavated in Yakutia and radiocarbon dated at least 62,400 years old. It became extinct about 13,000 years ago. Taxonomy ''Felis spelaea'' was the scientific name used by Georg Augus ...
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Late Pleistocene
The Late Pleistocene is an unofficial Age (geology), age in the international geologic timescale in chronostratigraphy, also known as Upper Pleistocene from a Stratigraphy, stratigraphic perspective. It is intended to be the fourth division of the Pleistocene Epoch within the ongoing Quaternary Period. It is currently defined as the time between c. 129,000 and c. 11,700 years ago. The Late Pleistocene equates to the proposed Tarantian Age of the geologic time scale, preceded by the officially ratified Chibanian (formerly known as Middle Pleistocene) and succeeded by the officially ratified Greenlandian. The estimated beginning of the Tarantian is the start of the Eemian interglacial period (Marine Isotope Stage 5). It is held to end with the termination of the Younger Dryas, some 10th millennium BC, 11,700 years ago when the Holocene Epoch began. The term Upper Pleistocene is currently in use as a provisional or "quasi-formal" designation by the International Union of Geological ...
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Mitochondrial
A mitochondrion (; ) is an organelle found in the cells of most Eukaryotes, such as animals, plants and fungi. Mitochondria have a double membrane structure and use aerobic respiration to generate adenosine triphosphate (ATP), which is used throughout the cell as a source of chemical energy. They were discovered by Albert von Kölliker in 1857 in the voluntary muscles of insects. The term ''mitochondrion'' was coined by Carl Benda in 1898. The mitochondrion is popularly nicknamed the "powerhouse of the cell", a phrase coined by Philip Siekevitz in a 1957 article of the same name. Some cells in some multicellular organisms lack mitochondria (for example, mature mammalian red blood cells). A large number of unicellular organisms, such as microsporidia, parabasalids and diplomonads, have reduced or transformed their mitochondria into other structures. One eukaryote, ''Monocercomonoides'', is known to have completely lost its mitochondria, and one multicellular organism, ...
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Subspecies
In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species have subspecies, but for those that do there must be at least two. Subspecies is abbreviated subsp. or ssp. and the singular and plural forms are the same ("the subspecies is" or "the subspecies are"). In zoology, under the International Code of Zoological Nomenclature, the subspecies is the only taxonomic rank below that of species that can receive a name. In botany and mycology, under the International Code of Nomenclature for algae, fungi, and plants, other infraspecific ranks, such as variety, may be named. In bacteriology and virology, under standard bacterial nomenclature and virus nomenclature, there are recommendations but not strict requirements for recognizing other important infraspecific ranks. A taxonomist decides whether ...
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Würm Glaciation
The Würm glaciation or Würm stage (german: Würm-Kaltzeit or ''Würm-Glazial'', colloquially often also ''Würmeiszeit'' or ''Würmzeit''; cf. ice age), usually referred to in the literature as the Würm (often spelled "Wurm"), was the last glacial period in the Alpine region. It is the youngest of the major glaciations of the region that extended beyond the Alps themselves. Like most of the other ice ages of the Pleistocene epoch, it is named after a river, in this case the Würm in Bavaria, a tributary of the Amper. The Würm ice age can be dated to about 115,000 to 11,700 years ago, but sources differ about the dates, depending on whether the long transition phases between the glacials and interglacials (warmer periods) are allocated to one or other of those periods. The average annual temperatures during the Würm ice age in the Alpine Foreland were below −3 °C (today +7 °C). That has been determined from changes in the vegetation (pollen analysis), as well a ...
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Excavation (archaeology)
In archaeology, excavation is the exposure, processing and recording of archaeological remains. An excavation site or "dig" is the area being studied. These locations range from one to several areas at a time during a project and can be conducted over a few weeks to several years. Excavation involves the recovery of several types of data from a site. This data includes artifacts (portable objects made or modified by humans), features (non-portable modifications to the site itself such as post molds, burials, and hearths), ecofacts (evidence of human activity through organic remains such as animal bones, pollen, or charcoal), and archaeological context (relationships among the other types of data).Kelly&Thomas (2011). ''Archaeology: down to earth'' (4th ed.). Belmont, Calif.: Wadsworth, Cengage Learning. Before excavating, the presence or absence of archaeological remains can often be suggested by, non-intrusive remote sensing, such as ground-penetrating radar. Basic informat ...
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Scientific Name
In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, although they can be based on words from other languages. Such a name is called a binomial name (which may be shortened to just "binomial"), a binomen, name or a scientific name; more informally it is also historically called a Latin name. The first part of the name – the '' generic name'' – identifies the genus to which the species belongs, whereas the second part – the specific name or specific epithet – distinguishes the species within the genus. For example, modern humans belong to the genus ''Homo'' and within this genus to the species ''Homo sapiens''. ''Tyrannosaurus rex'' is likely the most widely known binomial. The ''formal'' introduction of this system of naming species is credit ...
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Panthera Leo Spelaea MHNT
''Panthera'' is a genus within the family Felidae that was named and described by Lorenz Oken in 1816 who placed all the spotted cats in this group. Reginald Innes Pocock revised the classification of this genus in 1916 as comprising the tiger (''P. tigris''), lion (''P. leo''), jaguar (''P. onca''), and leopard (''P. pardus'') on the basis of common cranial features. Results of genetic analysis indicate that the snow leopard (formerly ''Uncia uncia'') also belongs to the genus ''Panthera'' (''P. uncia''), a classification that was accepted by IUCN Red List assessors in 2008. The tiger, lion, jaguar and leopard are the only cat species with anatomical structures that enable them to roar; the snow leopard cannot. The primary reason for this was formerly assumed to be the incomplete ossification of the hyoid bone. However, new studies show the ability to roar is due to other morphological features, especially of the larynx. Etymology The word ''panther'' derives from classical ...
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Radiocarbon Date
Radiocarbon dating (also referred to as carbon dating or carbon-14 dating) is a method for determining the age of an object containing organic material by using the properties of radiocarbon, a radioactive isotope of carbon. The method was developed in the late 1940s at the University of Chicago by Willard Libby. It is based on the fact that radiocarbon () is constantly being created in the Earth's atmosphere by the interaction of cosmic rays with atmospheric nitrogen. The resulting combines with atmospheric oxygen to form radioactive carbon dioxide, which is incorporated into plants by photosynthesis; animals then acquire by eating the plants. When the animal or plant dies, it stops exchanging carbon with its environment, and thereafter the amount of it contains begins to decrease as the undergoes radioactive decay. Measuring the amount of in a sample from a dead plant or animal, such as a piece of wood or a fragment of bone, provides information that can be used to calcu ...
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Yakutia
Sakha, officially the Republic of Sakha (Yakutia),, is the largest republic of Russia, located in the Russian Far East, along the Arctic Ocean, with a population of roughly 1 million. Sakha comprises half of the area of its governing Far Eastern Federal District, and is the world's largest country subdivision, covering over 3,083,523 square kilometers (1,190,555 sq mi). ''Sakha'' following regular sound changes in the course of development of the Yakut language) as the Evenk and Yukaghir exonyms for the Yakuts. It is pronounced as ''Haka'' by the Dolgans, whose language is either a dialect or a close relative of the Yakut language.Victor P. Krivonogov, "The Dolgans’Ethnic Identity and Language Processes." ''Journal of Siberian Federal University'', Humanities & Social Sciences 6 (2013 6) 870–888. Geography * ''Borders'': ** ''internal'': Chukotka Autonomous Okrug (660 km)(E), Magadan Oblast (1520 km)(E/SE), Khabarovsk Krai (2130 km)(SE), Amur Oblast (S ...
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Proceedings Of The National Academy Of Sciences Of The United States Of America
''Proceedings of the National Academy of Sciences of the United States of America'' (often abbreviated ''PNAS'' or ''PNAS USA'') is a peer-reviewed multidisciplinary scientific journal. It is the official journal of the National Academy of Sciences, published since 1915, and publishes original research, scientific reviews, commentaries, and letters. According to ''Journal Citation Reports'', the journal has a 2021 impact factor of 12.779. ''PNAS'' is the second most cited scientific journal, with more than 1.9 million cumulative citations from 2008 to 2018. In the mass media, ''PNAS'' has been described variously as "prestigious", "sedate", "renowned" and "high impact". ''PNAS'' is a delayed open access journal, with an embargo period of six months that can be bypassed for an author fee ( hybrid open access). Since September 2017, open access articles are published under a Creative Commons license. Since January 2019, ''PNAS'' has been online-only, although print issues are ava ...
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Nuclear Genome
Nuclear DNA (nDNA), or nuclear deoxyribonucleic acid, is the DNA contained within each cell nucleus of a eukaryotic organism. It encodes for the majority of the genome in eukaryotes, with mitochondrial DNA and plastid DNA coding for the rest. It adheres to Mendelian inheritance, with information coming from two parents, one male and one female—rather than matrilineally (through the mother) as in mitochondrial DNA. Structure Nuclear DNA is a nucleic acid, a polymeric biomolecule or biopolymer, found in the nucleus of eukaryotic cells. Its structure is a double helix, with two strands wound around each other, a structure first described by Francis Crick and James D. Watson (1953) using data collected by Rosalind Franklin. Each strand is a long polymer chain of repeating nucleotides. Each nucleotide is composed of a five-carbon sugar, a phosphate group, and an organic base. Nucleotides are distinguished by their bases: purines, large bases that include adenine and guanine; and pyr ...
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Molecular Ecology
Molecular ecology is a field of evolutionary biology that is concerned with applying molecular population genetics, molecular phylogenetics, and more recently genomics to traditional ecological questions (e.g., species diagnosis, conservation and assessment of biodiversity, species-area relationships, and many questions in behavioral ecology). It is virtually synonymous with the field of "Ecological Genetics" as pioneered by Theodosius Dobzhansky, E. B. Ford, Godfrey M. Hewitt, and others. These fields are united in their attempt to study genetic-based questions "out in the field" as opposed to the laboratory. Molecular ecology is related to the field of conservation genetics. Methods frequently include using microsatellites to determine gene flow and hybridization between populations. The development of molecular ecology is also closely related to the use of DNA microarrays, which allows for the simultaneous analysis of the expression of thousands of different genes. Quanti ...
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